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The ongoing lower level of species extinction intensity occurring between episodes of mass extinctions. Extinction is a normal part of the history of life. Out of every 100 species that have ever lived on Earth, 99 are no longer in existence.…

The adaptation of an animal or plant through breeding in captivity to a life intimately associated with and advantageous to humans. Domesticated plants and animals differ genetically and morphologically from their wild counterparts, often in ways…

Typically, an aquatic vertebrate with scales, gills, and paired fins. Fishes (Fig. 1) are found throughout the world's waterways, from saltwater oceans to freshwater streams and ponds. About 35,000 species of fishes have been described to date. Many…

The use of pollen and spore evidence in legal cases. In its broader application, the field of forensic palynology also includes legal information derived from the analysis of a broad range of microscopic organisms that can be found in both fresh and…

A plant that has altered genetic material (DNA) through recombinant DNA (rDNA) manipulation. Genetic engineering of plants allows for very specific genetic changes to be made to a plant's deoxyribonucleic acid (DNA), as opposed to the random…

A commercially grown plant that has altered genetic material through artificial recombinant manipulation. Genetic modification is the latest scientific tool for developing improved crop varieties. Such genetically modified (GM) crops can help to…

A biological organism whose genetic material has been manipulated artificially through genetic or genomic editing techniques. In contrast to the long history of artificial selection (domestication), in which humans selectively breed organisms to…

A space-traversing unmanned vehicle which contains the means for controlling its flight path and that delivers an explosive warhead with high accuracy. Various kinds of missiles are guided in the sense that they can aerodynamically adjust after…

A subdiscipline of paleobiology that investigates the processes of preservation and their influence on information in the fossil record. Taphonomy is the study of fossilization and considers the processes at work in the incorporation of living…

A plant, Musa textilis, of the banana family that is valuable for its hard fiber (also known as Manila hemp). Abaca (Musa textilis) is a flowering plant native to the Philippines and Borneo. Taxonomically, it is assigned to the order Zingiberales.…

A major plant hormone that plays an important role in plant growth and development. Abscisic acid (ABA), with a chemical formula of C15H20O4 (Fig. 1), is one of the most significant hormones found in plants. It is responsible for regulating various…

Mechanized systems of food and fiber production used in agriculture. The mechanized systems comprising agricultural machinery are utilized in numerous aspects of agriculture, including the initial tillage of the soil (Fig. 1), planting, cultural…

An informal assemblage of predominantly aquatic eukaryotes that carry out oxygen-evolving photosynthesis, but lack specialized water-conducting and food-conducting tissues. Algae (Fig. 1) are eukaryotic organisms (having an organized nucleus).…

A small, tropical, evergreen tree (Pimenta dioica) and its dried, unripe fruit (berry). Allspice (Pimenta dioica; alternatively, P. officinalis) [see illustration] is a member of the myrtle family (Myrtaceae). Allspice, which is also known as…

Most commonly, a generic name for all fossil resins, although it has been restricted by some to refer only to succinite, the mineralogical species of fossil resin that includes most of the Baltic Coast deposits. The time necessary to transform resin…

The use of live nonhuman animals for scientific research and experimentation. Animal testing has been a cornerstone of biomedical research, having contributed greatly to the understanding of human biology, to the unraveling of the complex underlying…

A genus of fossil birds in the extinct order Archaeopterygiformes, characterized by flight feathers like those of modern birds. Archaeopteryx is generally accepted as the oldest known fossil bird. It represents a unique snapshot of evolution; most…

The application of chemical techniques to the study of archeological materials in order to elucidate aspects of the human past. The materials used in archeological chemistry might be anthropogenic (for example, pottery or metal) or natural (from…

Any vegetable that is prevalent in Asian cuisines, but less well known in Western diets. Asian vegetables (Fig. 1), also referred to as Asian greens or oriental vegetables, are very important in Asian countries, but are not as popular in North…

Any of a class of plant growth substances (phytohormones or plant hormones) that have an essential role in coordination of many growth and developmental processes during the life cycle of plants, especially the ability to increase the rate of shoot…

A class of nonflagellate unicellular algae, commonly called diatoms, with boxlike silicified walls. Members of the algal class Bacillariophyceae are commonly termed diatoms (Fig. 1). Diatoms range in maximum dimension from 4 μm (0.00004 in.) to more…

Any of several leguminous plants, or their seeds, long utilized as food by humans or livestock. There are more than 40,000 varieties of beans, with only a small number of these being mass-produced for human or animal consumption. In particular,…

The drug as well as the plant known as deadly nightshade (Atropa belladonna). The belladonna plant (Atropa belladonna; see illustration) belongs to the nightshade family, Solanaceae, in the order Solanales. The plant is a coarse and poisonous…

The study of skeletal remains from archeological sites by biological (or physical) anthropologists. Bioarcheology differs in several ways from traditional skeletal research (see illustration). Previous work focused on individual case studies (for…

Fungi that exhibit the natural phenomenon of bioluminescence (the ability of an organism to emit visible light). Bioluminescence is the emission of photons of visible light by living organisms via the chemical reaction of molecular oxygen (O2) with…

A garden for the culture of plants collected chiefly for scientific, educational, and esthetic purposes. A botanical garden (see illustration) is an establishment in which various plants, including trees, are collected, cultivated, and grown for…

The branch of biological science that embraces the study of plants and plant life. Botany is the scientific study of plants (see illustration), encompassing investigations of plant anatomy, physiology, genetics, phylogeny, ecology, pathology, and…

A division that consists of small and relatively simple plants commonly known as mosses, peat mosses, granite mosses, liverworts, and hornworts. Members of the division Bryophyta are referred to as bryophytes (see illustration). These nonvascular…

A genus comprising yeasts that cause a variety of human diseases referred to as candidiasis. All Candida species are single-celled yeasts (fungal organisms) that are ovoid in shape and 3–5 μm in size, and they multiply by a process called budding.…

An order of flowering plants (angiosperms) including the mustard and caper families. The plant order Capparales contains various families of eurosid dicotyledons. Most notably, mustards, capers, and various ornamental plants (see illustration) are…

A member of a class of common pigments, typically of yellow, orange, or red hues, that serve in both light absorption and the protection against too much light in photosynthesis as well as in the vision process. Carotenoids comprise a class of…

A medium-sized, spreading evergreen tree (Anacardium occidentale), which is grown widely in the tropics for its kidney-shaped edible nuts (also called cashews) and the resinous oil in the shells of the nuts. Cashew trees, which are native to Brazil,…

A coarse, erect, annual herb plant, Ricinus communis, belonging to the spurge family (Euphorbiaceae). The castor plant (Ricinus communis) is a member of the spurge family (Euphorbiaceae) in the order Malpighiales. The plant varies greatly in height…

A class in the phylum Mollusca that is exclusively marine and includes squids, octopuses, and cuttlefishes. The molluscan class Cephalopoda is one of the major taxonomic groupings of marine animals and contains both extant members—squids (Fig. 1),…

Any member of the grass family (Poaceae or Gramineae) that produces edible grains usable as food by humans and livestock. Cereals are important cultivated crops. Common cereals include rice, wheat, barley, oats, maize (corn), sorghum, rye, and…

An order of the class Reptilia, including the turtles, terrapins, and tortoises; also termed Testudines. The order Chelonia (often referred to as Testudines) comprises one of the oldest groups of reptiles on Earth. Members of the Chelonia…

A type of sweet or sour fruit of the Prunus genus, as well as the tree that bears the fruit. Cherry trees belong to the Prunus genus in the order Rosales, and the sweet and sour fruits of these trees are called cherries (Fig. 1). There are…

An evergreen shrub or small tree, particularly Cinnamomum verum, and the spice obtained from the plant's bark. The cinnamon plant (see illustration) is a member of the laurel family (Lauraceae) in the order Laurales. Native to Sri Lanka, the…

A large palm, Cocos nucifera, widely grown throughout the tropics and valuable for its fruit and fiber. The coconut (Cocos nucifera) belongs to the plant family Arecaceae. The name coconut can refer to the tree and its fruit. Usually found near the…

Any of various tropical trees belonging to the genus Cola. Members of the genus Cola (family Malvaceae, order Malvales), which are native to tropical Africa, are called cola trees (see illustration). In particular, two species, C. acuminata and C.…

Any living or fossil member of the plant division Pinophyta, consisting chiefly of woody trees or shrubs that produce cones and evergreen needles. Conifers are predominantly cone-bearing, evergreen trees and shrubs belonging to the order Pinales, as…

A fiber obtained from the cotton plant Gossypium. The cotton plant (genus Gossypium, order Malvales; Fig. 1) has been used by humans for more than 5000 years. The fiber (cotton) obtained from this plant is the most widely used natural fiber because…

An unharvested crop grown to improve soil quality or enhance pest management. Increasing numbers of farmers plant cover crops as a means of conserving soil, enhancing subsequent production, and reducing off-farm inputs. The sustainable agriculture…

A woody higher plant that regularly loses its leaves at the end of each growing season. Deciduous plants drop (shed) their leaves seasonally, most commonly during the fall (autumn). In general, the dropping of leaves occurs at the inception of an…

The deliberate conversion of forest to nonforest land that results in the long-term reduction of tree canopy below a minimum level. Deforestation, also known as land clearing, permanently eliminates forests to utilize the land for other purposes…

The biological study of insects that have a direct influence on humanity. Economic entomology is a subdiscipline of entomology (the biological science that deals with the investigation of insects) and focuses on the effects of insects on human…

A System that locates suitably equipped users by means of measurements made by electronic sensors. Typical measurements include distance or bearing from a vehicle to a known point, or direct determination of the present position of the vehicle in a…

A major clade or natural group of plants, comprising the land plants. The members of the Embryobionta or Embryophyta are commonly termed embryobionts or embryophytes, and they constitute the land plants. Embryophytes form a well-marked clade (a…

A volatile, fragrant oil obtained from plants. Essential oils are distinguished from those known as fixed oils, which are mainly triglycerides of fatty acids. Essential oils have been obtained from thousands of plants (Fig. 1) and are designated and…

A former order of rosid flowering plants, division Magnoliophyta (Angiospermae), in the class Magnoliopsida (dicotyledons). The order Euphorbiales consisted of the large family Euphorbiaceae (with about 7500 species) and 3 small satellite families…

An active and intentional act or practice of putting to death, in a relatively painless way, of persons suffering severely from medical conditions that are incurable, untreatable, or irreversible. Euthanasia is an act by a person, usually a…

A plant that retains its green foliage throughout the year. An evergreen plant, or simply evergreen, is a tree, shrub, or plant whose leaves are not shed in the winter and instead remain green year-round. In contrast, deciduous trees and plants…

The test of a hypothesis under controlled conditions. The experiment is one of the distinctive tools of the scientist. It enables the scientist to put questions to nature and receive answers. These answers lead to new problems whose solutions…

The death and disappearance of a species. The extinction of a species usually represents an end point in a long series of population extinctions. During the extinction process, unique evolutionary history is lost at every stage, but the death of the…

Plants grown and harvested for food, feed, or industrial purposes. Farm crops may be roughly classified as follows: (1) food crops—the bread grains [wheat (see illustration) and rye], rice, sugar crops (sugarbeets and sugarcane), potatoes, and dry…

An enzymatic transformation of organic substrates, especially carbohydrates, generally accompanied by the evolution of a gas. Fermentation is a chemical transformation of organic substrates, especially carbohydrates, to alcohol or organic acids,…

Materials added to the soil, or applied directly to crop foliage, to supply elements needed for plant nutrition, and the management of this nutrient application to sustain maximum plant growth and yield while minimizing environmental impacts of…

A plant grown for its fibrous material, which is used for many commercial purposes and for home industry. Many plants are grown as fiber crops because of their content or yield of fibrous material (see illustration), which is used in the manufacture…

The segment of horticulture concerned with the commercial production, marketing, and sale of plants grown and used for decorative or display purposes. Floriculture is a branch of horticulture, that is, the art and science of growing plants,…

The application of different types of energy and techniques involving equipment and apparatuses to transform agricultural products of plant and animal origin or seafood into food ingredients, semifinished commodities, and finished ready-to-eat…

The study of the physical, chemical, and biological properties of foods as well as the sensory, nutritional, storage properties, and safety of foods. Food properties are influenced by growing, harvesting, and slaughtering practices, preservation and…

Any of the grasses and legumes that make up grasslands and are consumed by grazing livestock. Grasslands represent an ancient renewable natural resource. In general, grasslands form 25% of the world's vegetation and occupy the largest area of any…

The natural medium for growth of tree roots and associated forest vegetation. Proper forest growth and management depend on many factors, including the quality of the forest soil (see illustration). In general, weathering of underlying rock,…

The science, art, and practice of managing and using the natural resources that occur on and in association with forested lands. Forestry is the management of forested lands (see illustration). It involves the planning and implementation of…

A record of earlier life buried in rock. Originally meaning any distinctive object that has been dug up (from Latin fodio, dig), the term “fossil” soon came to refer particularly to things resembling animals and plants. These objects were known…

A hypersensitivity or altered reactivity caused by fungal allergens. More than 20% of the population in industrialized countries suffers from allergic disease, and diverse allergens (substances foreign to the body that cause allergy) associated with…

A hardy perennial, Allium sativum, of Asiatic origin and belonging to the plant order Asparagales. Garlic is grown for its pungent bulbs (see illustration). Segments of these are used primarily for seasoning. Europeans have grown garlic for probably…

Soil ingestion by animals. Grazing animals such as sheep and cattle ingest varying amounts of soil when they graze herbage contaminated with it.Pastures become contaminated with soil when livestock walk across the herbage, particularly in wet…

A member of a family of diterpenoid carboxylic acids that are notable for being plant hormones. Gibberellins are diterpenoid carboxylic acids. Most of them possess ent-gibberellane or ent-20-norgibberellane carbon (C) skeletons. Gibberellins occur…

An important spice or condiment, and the plant (Zingiber officinale) from which it is obtained. The ginger plant, Zingiber officinale, is a member of the ginger family Zingiberaceae and is a native of southeastern Asia. The plant is an erect…

The common name for a group of perennial herbs in the genus Panax cultivated for the aromatic root of the plant, which is used medicinally in China. Herbal members of the genus Panax belong to the aralia family (Araliaceae) and are native to…

Any of the crop plants belonging to the grass family (Poaceae or Gra­mineae) that are generally grown for their edible starchy seeds. Grain crops are often referred to as cereal crops and include wheat, rice, corn (maize), barley (see illustration),…

The common name for plants of the genus Vitis characterized by climbing stems and grown for the edible, pulpy berries. Grapes (genus Vitis) comprise one of the world's most significant and economically important crops. The common grapevine is…

An evergreen tree (Citrus paradisi) and its edible citrus fruit. Grapefruit (Citrus paradisi) is a subtropical evergreen tree belonging to the order Sapindales. It has a well-rounded top and is cultivated for its large and globose citrus fruit. The…

Any member of the family Poaceae (Gramineae) cultivated as forage and grain for consumption. Grasses are arguably the most useful of all the plants that cover the Earth. The cereal grass crops [wheat (Fig. 1), rice, corn (maize), rye, barley, oats,…

A morphologically distinct cell that is found in the epidermis of terrestrial plant shoots and that functions in regulating stoma size. Guard cells are paired, crescent-shaped epidermal cells that regulate water loss and gas exchange in terrestrial…

Optical instruments that establish an optical line or axis for the purpose of aiming a weapon. The axis includes the observer's eye, a suitable mark in the instrument, and the target. For a pistol, the traditional iron sights employ a rear notch and…

A class of lower green plants called liverworts that belong to the division Bryophyta. The class Hepaticopsida, also called Marchantiopsida, comprises the liverworts, which are nonvascular land plants. Although there is a great diversity of external…

An intensively managed plant cultivated for food or for esthetic purposes. Plant agriculture is divided traditionally into the fields of agronomy (herbaceous field crops, mainly grains, forages, oilseeds, and fiber crops), forestry (forest trees and…

The collective techniques for supplying nutrients and water directly to the roots of plants, without soil or other media. Hydroponics is concerned with the cultivation and growing of plants without soil or other media. Hydroponic systems (Fig. 1)…

A tentative supposition with regard to an unknown state of affairs, the truth of which is thereupon subject to investigation by any available method, either by logical deduction of consequences which may be checked against what is known, or by…

Any of several low, perennial, tropical South American shrubs or half-shrubs in the family Rubiaceae. Ipecac plants (also termed ipecacuanha plants) are native to the tropical forests of Brazil and Colombia. Several species are used, but the dried…

The artificial application of water to the soil to produce plant growth. Irrigation also cools the soil and atmosphere, making the environment favorable for plant growth. The use of some form of irrigation (Fig. 1) is well documented throughout the…

Intentional generation of interfering signals in the electromagnetic spectrum by powerful transmitters as a countermeasure intended to block a communication, radar, or infrared system or to impair its effectiveness appreciably. Radio broadcasts or…

The common name for a member of the agnathan fishes. The fish superclass Agnatha (subphylum Vertebrata, phylum Chordata) comprises the jawless vertebrates (agnathans). Members include cyclostomes (modern lampreys and hagfishes), various extinct…

The removal of water from the surface of the land and the control of the shallow ground water table improves the soil as a medium for plant growth. The sources of excess water may be precipitation, snowmelt, irrigation water, overland flow or…

An order of flowering plants composed of seven eumagnoliid families of tropical tree and shrub species. The plant order Laurales, with approximately 2500–2800 species, contains numerous trees and shrubs having ecological significance. Members of the…

Citrus limon, a small evergreen tree belonging to the order Sapindales that is cultivated for its fruit. In commercial terms, the lemon (Citrus limon; order Sapindales) is the most important of the acid citrus fruits (see illustration). It probably…

Lama glama, a member of the camel family (Camelidae) found only in South America. The llama (see illustration) is an artiodactyl, or even-toed ungulate, belonging to the mammalian family Camelidae. It has two toes on each foot. The llama is a…

Solvent removal from the frozen state by sublimation; commonly referred to as freeze-drying. In lyophilization, the dried material, a porous solid, is sealed under vacuum or inert-gas (liquid nitrogen) atmosphere and retains its physical and…

One of the largest orders of the rosid eudicotyledons. The order Malpighiales is an important flowering plant group with a worldwide distribution. Depending on the classification, it contains approximately 35 to 40 families, totaling about 16,000…

An order of flowering plants in the core eudicots. The plant order Malvales consists of 10 families and approximately 6000 species (see illustration). The circumscription of the order Malvales has been altered greatly as a result of deoxyribonucleic…

A lineage of true elephants in the extinct genus Mammuthus that occupied Eurasia and North America in the late Cenozoic Era. Mammoths (genus Mammuthus) are extinct elephants assigned to the mammalian order Proboscidea (Fig. 1). They inhabited…

A psychoactive narcotic drug obtained from the dried leaves, stems, seeds, and flowering tops of the hemp plant, Cannabis sativa. Cannabis sativa (Fig. 1), commonly called cannabis, is the source of marijuana, which is one of the most widely used…

The widespread, catastrophic loss of many species in a relatively short period of geological time. Earth has supported life for approximately 3.5 billion years, and it is normal for species to become extinct and for new species to arise through…

The body of knowledge that concerns the fundamental nature, function, development, and use of formal models in science and technology. In its most general sense, a model is a proxy. A model is one entity used to represent some other entity for some…

A major phylum of the animal kingdom comprising a diverse group of unsegmented, soft-bodied invertebrates. The phylum Mollusca is one of the most significant groupings of invertebrate animals. Members of the Mollusca, termed mollusks or molluscs,…

A mammalian order containing the egg-laying platypus and echidnas. Members of the order Monotremata (monotremes) are egg-laying mammals belonging to the subclass Prototheria (in some alternative classifications, the monotremes have been assigned to…

A macroscopic fungus with a fruiting body. It has been estimated that 1.5 million to 5 million species of fungi exist on Earth. Yet, only about 150,000 species have been described to date. Approximately 50,000 fungal species are considered…

Weapons that arm warships. Of the wide variety of such weapons, three important categories are exclusively naval: antiship missiles, torpedoes, and mines. Other weapons all correspond to land-based types, but submarine-launched ballistic missiles…