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A diverse group of early jawed fishes from the Paleozoic Era, usually classified as a distinct class of vertebrates. Most acanthodians have tiny body scales with a cross section like that of an onion, the layers reflecting incremental scale growth.…

An order of actinopterygian (ray-finned) fishes represented by the sturgeons and paddlefishes. The order Acipenseriformes, the only extant order of about 10–12 orders of chondrosteans (subclass Chondrostei), probably originated from palaeonisciform…

An order of inarticulated brachiopods. Together with Lingulida and Siphonotretida, Acrotretida is one of the three orders of inarticulated brachiopods currently classified in class Lingulata in subphylum Linguliformea (phylum Brachiopoda).…

A class of teleostom fishes commonly known as the ray-finned fishes, which are distinguished by the structure of the paired fins that are supported by dermal rays. The class Actinopterygii comprises the subclasses Cladistia [Polypteriformes (bichirs…

An order of extremely elongate, limbless fossil amphibians in the subclass Lepospondyli from Permo-Carboniferous rocks of North America and Europe. The order Aistopoda (also spelled Aïstopoda) consists of extremely elongate, limbless fossil…

Most commonly, a generic name for all fossil resins, although it has been restricted by some to refer only to succinite, the mineralogical species of fossil resin that includes most of the Baltic Coast deposits. Resins generally are complex mixtures…

An extinct subclass of Cephalopoda (phylum Mollusca). Besides the Ammonoidea (chambered ammonoids), the other subclasses of Cephalopoda are Nautiloidea (chambered nautiloids) and Coleoidea (squid, cuttlefish, octopus, and extinct belemnites), both…

One of four classes (the others are Reptilia, Aves, and Mammalia) composing the superclass Tetrapoda of the subphylum Vertebrata. The living amphibians number over 4700 species and are classified in three orders: Salientia or Anura (frogs and toads,…

Formerly, a group of reptiles that was recognized in rank-based classifications. Amniotes were once divided into subclasses, based on the patterns of temporal fenestrae (openings in the skull roof posterior to the openings for the eyes). Subclass…

A group of fossil jawless fishes, roughly similar in appearance to extant lampreys, known from Early Silurian–Early Devonian deposits (about 410–430 million years ago) that represent marine, coastal environments. Anaspid fossils are known…

The phylum comprising the multisegmented, invertebrate wormlike animals, of which the most numerous are the marine bristle worms and the most familiar the terrestrial earthworms. The Annelida (meaning little annuli or rings) include the Polychaeta…

An order of Paleozoic tetrapods that arose Carboniferous (Mississippian) (345 million years ago) and lasted until the Early Triassic (about 245 MYA). The term anthracosaurs (meaning “coal reptiles”) encompasses a range of genera that may not all be…

The smallest order of the class Amphibia, consisting of the wormlike caecilians. The order Apoda is sometimes called Gymnophiona. Its members are known commonly as caecilians, which are wormlike, legless amphibians with indistinct or even hidden…

An order of Paleozoic diapsid reptiles including the families Petrolacosauridae and Araeoscelidae. Members of these families resemble primitive modern lizards, such as the green iguana, in size and most body proportions, but are distinguished by…

An extinct group of mainly Lower Cambrian marine sponges which, although lacking spicules, possessed an intricate, highly porous skeleton of calcite. It was probably a monophyletic group; that is, all representatives were derived from a single…

An order of extinct, free-sporing plants (cryptogams) that lived during the Late Devonian through Early Mississippian. The order includes the genera Archaeopteris (which has been extensively investigated), Actinoxylon, Actinopodium, Svalbardia,…

A genus of fossil birds in the extinct order Archaeopterygiformes, characterized by flight feathers like those of modern birds. Archaeopteryx is generally accepted as the oldest known fossil bird. It represents a unique snapshot of evolution; most…

The most diverse group of early Tertiary mammals are the archaic ungulates (condylarths). Although closely related, they are not a monophyletic group but, along with various extinct and extant mammals, form the taxon Ungulata (Fig. 1). They are…

The establishment of the temporal sequence of prehistoric events. An event can be defined as a duration of time during which certain features came together to form a set, for example, when certain materials were formed into an object (manufacturing…

A diversified group of diapsid reptiles, comprising dinosaurs (and their descendants, birds), crocodylians, pterosaurs (flying reptiles), and various forms closely related to these groups. In the past, the primitive archosaurs from the Triassic…

A heavily armored mammal in the order Cingulata. The order Cingulata (formerly Edentata or Xenarthra) comprises the armadillos, which are divided into two extant families: Chlamyphoridae and Dasypodidae. These creatures are characterized by bony…

The largest phylum in the animal kingdom, including the well-known insects, spiders, ticks, and crustaceans, as well as many smaller, less well-known groups. Members of the phylum Arthropoda, that is, arthropods, are the dominant metazoans…

The only surviving subclass of the echinoderm class Crinoidea. The Articulata include all living members of the echinoderm class Crinoidea and are generally regarded as the post-Paleozoic crinoids. Crinoids are exclusively marine organisms, as are…

An order comprising the even-toed ungulates (hoofed mammals). There are two main radiations: the predominantly omnivorous Bunodontia, including suoids (such as pigs, peccaries, and hippos); and the more herbivorous Selenodontia, including camels and…

A small order of specialized holostean fishes which are first recorded from Middle Jurassic deposits of Europe. Later records indicate that they were a very successful group in terms of geographic range, and probably had a worldwide distribution in…

A relatively small order of extinct South American ungulates in the infraclass Eutheria, known from the late Paleocene to the late Miocene. Members of the order Astrapotheria are customarily divided into two suborders: the late Paleocene–Eocene…

Any member of an adaptively diverse group of extinct hominin species that inhabited Africa between approximately 4.2 million years ago (MYA) and 1.2 MYA. The term australopith is used in preference to the term australopithecine, which is sometimes…

The class of animals consisting of the birds. Birds (Fig. 1) are vertebrate animals comprising the class Aves. (In some classification schemes, the term Neornithes may also be used when referring to modern birds.) Primarily adapted for flying, the…

The ongoing lower level of species extinction intensity occurring between episodes of mass extinctions. Extinction is a normal part of the history of life. Out of every 100 species that have ever lived on Earth, 99 are no longer in existence.…

An elasmobranch subdivision of cartilaginous fishes known collectively as the rays, skates, or batoids. Batoidea is one of the two subdivisions of the subclass Elasmobranchii (class Chondrichthyes) and consists of 4 extant orders, 17–20 families,…

An order of extinct 10-armed cephalopod mollusks with an internal shell. The order Belemnitida belongs to the subclass Coleoidea in the class Cephalopoda (phylum Mollusca). Constituent members, which are all extinct, are known as belemnites (Fig. 1)…

An extinct group of epifaunal bivalves that constructed reefs during the late Mesozoic. The valves of rudists are coiled to cylindrical in external form, and the shapes and sizes of rudist shells are extreme relative to other members of the class…

One of the seven existing classes in the phylum Mollusca; sometimes known as Pelecypoda. All members of the Bivalvia (bivalves) are aquatic, living at all depths of the sea and in brackish waters and freshwaters. With about 25,000 living species…

A class of extinct Pelmatozoa in the subphylum Blastozoa which arose in the Ordovician and flourished in the Carboniferous and Permian. Blastoids did not survive the Permo-Triassic mass extinction event. Historically, two orders of the Blastoidea…

A phylum of solitary, exclusively marine, coelomate, bivalved animals, with both valves symmetrical about a median longitudinal plane. Brachiopods are typically attached to the substrate by a posteriorly located cuticle-covered stalk called a…

A phylum of sessile aquatic invertebrates (also called Polyzoa) that form colonies of zooids. Each zooid, in its basic form, has a lophophore of ciliated tentacles situated distally on an introvert, a looped gut with the mouth inside the lophophore…