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Determination of the relative or absolute age of materials or objects by measurement of the degree of racemization of the amino acids present. With the exception of glycine, the amino acids found in proteins can exist in two isomeric forms called d-…

An extinct subclass of Cephalopoda (phylum Mollusca). Besides the Ammonoidea (chambered ammonoids), the other subclasses of Cephalopoda are Nautiloidea (chambered nautiloids) and Coleoidea (squid, cuttlefish, octopus, and extinct belemnites), both…

A collective term for the classes Reptilia (reptiles), Aves (birds), and Mammalia (mammals) of the subphylum Vertebrata. The remaining vertebrates, including the several classes of fishes and the amphibians, are grouped together as the Anamnia.…

An order of Lobosia without protective coverings (tests). These protozoa range in size from about 4 micrometers to 0.08–0.12 in. (2–3 mm). Pellicles may be thin, as in Amoeba proteus, or thicker and less flexible, as in Thecamoeba verrucosa (Fig.…

One of four classes (the others are Reptilia, Aves, and Mammalia) composing the superclass Tetrapoda of the subphylum Vertebrata. The living amphibians number over 4700 species and are classified in three orders: Salientia or Anura (frogs and toads,…

Collectively, members of the class Amphibia are facing massive reductions in total numbers, and the overall biodiversity of amphibians across the globe is under threat. Scientists have determined that global climate change and habitat destruction…

A mineral with a framework structure in which all the aluminosilicate tetrahedral vertices are linked, thus allying it to the feldspars, feldspathoids, and zeolites. The structure is cubic, space group Ia3d, a = 1.371 mm. Its formula is…

A collective term, also referred to as Anamniota, for the vertebrate animals that lack an amnion in development, including the Agnatha, Chondrichthyes, Osteichthyes, and Amphibia. The amnion is a protective embryonic envelope that encloses the…

Formerly, a group of reptiles that was recognized in rank-based classifications. Amniotes were once divided into subclasses, based on the patterns of temporal fenestrae (openings in the skull roof posterior to the openings for the eyes). Subclass…

A group of fossil jawless fishes, roughly similar in appearance to extant lampreys, known from Early Silurian–Early Devonian deposits (about 410–430 million years ago) that represent marine, coastal environments. Anaspid fossils are known…

An extant order of the crustacean superorder Syncarida confined to freshwater and recorded from both surface and subterranean waters. Members of the order Anaspidacea (anaspidaceans) lack a carapace. The first thoracomere (thoracic somite) is…

A nesosilicate mineral, composition Al2SiO5, crystallizing in the orthorhombic system. It occurs commonly in large, nearly square prismatic crystals. The variety chiastolite has inclusions of dark-colored carbonaceous material arranged in a regular…

A plagioclase feldspar with composition Ab70An30 to Ab50An50 (Ab = NaAlSi3O8; An = CaAl2Si2O8). Andesine occurs primarily in igneous rocks, often in a glassy matrix as small, chemically zoned, lathlike crystals known as microlites. The rock types…

A typical volcanic rock erupted from a volcano associated with convergent plate boundaries. The process of subduction, which defines convergent plate boundaries, results in the sinking of oceanic lithosphere beneath either oceanic lithosphere or…

A mineral with the chemical composition PbSO4. Anglesite occurs in white or gray, orthorhombic, tabular or prismatic crystals or compact masses (see illustration). It is a common secondary mineral, usually formed by the oxidation of galena. Fracture…

The true eels, a large order of actinopterygian fishes, also known as the Apodes. The order Anguilliformes is related to Saccopharyngiformes (sackpharynx fishes), Elopiformes (tarpons), and Notacanthiformes (spiny eels and halosaurs); all members…

A mineral with the chemical composition CaSO4. Anhydrite occurs commonly in white and grayish granular masses, rarely in large, orthorhombic crystals (see illustration). Fracture is uneven and luster is pearly to vitreous. Hardness is 3–3.5 on Mohs…

Substances suitable for the nutrition of animals, including livestock and poultry feed and pet food. Animal feed includes plant and animal products and by-products as well as vitamin and mineral supplements. Livestock and poultry feed In formulating…

An animal's movement through the atmosphere sustained by aerodynamic reaction or other forces. Animal flight includes gliding and flapping flight. Gliding flight does not involve the generation of thrust, whereas flapping flight does generate thrust…

The classification rank comprising the animals. Animals are taxonomically assigned to the classification kingdom known as Animalia. (The other kingdoms for classification of organisms include Plantae, Fungi, Protista, and Monera; in certain…

An annual herb, Pimpinella anisum (family Apiaceae), whose fruit is used to provide food flavoring and to make a distilled oil. Anise is one of the earliest aromatics mentioned in literature. The anise plant (see illustration) is an annual herb that…

The carbonate mineral Ca(Fe,Mg)(CO3)2, also commonly containing some manganese. The mineral has hexagonal (rhombohedral) symmetry (see illustration) and has the cation-ordered structure of dolomite. The name is applied only to those species in which…

The phylum comprising the multisegmented, invertebrate wormlike animals, of which the most numerous are the marine bristle worms and the most familiar the terrestrial earthworms. The Annelida (meaning little annuli or rings) include the Polychaeta…

The calcium-rich plagioclase feldspar with composition Ab0An100 to Ab10An90 (Ab = NaAlSi3O8; An = CaAl2Si2O8), occurring in olivine-rich igneous rocks and rare volcanic ejecta (for example, at Mount Vesuvius, Italy, and Miyakejima, Japan). Hardness…

The name usually given to alkali feldspars which have a chemical composition ranging from Or40Ab60 to Or10Ab90± up to approximately 20 mole % An (Or, Ab, An = KAlSi3O8, NaAlSi3O8, CaAl2Si2O8) and which deviate in one way or another from monoclinic…

A rock composed of 90 vol % or more of plagioclase feldspar. Strictly, the rock is composed entirely of crystals discernible with the eye, but some finely crystalline examples from the Moon have been called anorthosite or anorthositic breccia. Two…

An order of branchiopod crustaceans, known as fairy shrimps and, in some cases, brine shrimps (Artemia). Members of the order Anostraca (anostracans) range up to about 100 mm (4 in.) in length but usually are much smaller. The trunk consists of…

Oxygen-depleted regions in marine environments. The dynamic steady state between oxygen supply and consumption determines the oxygen concentration. In regions where the rate of consumption equals or exceeds the rate of supply, seawater becomes…

An order of birds comprising two families, the screamers (Anhimidae) of South America and the worldwide waterfowl (Anatidae). They are closely related to the Galliformes (fowls), with the screamers being a rather intermediate group. The waterfowl…

The Antarctic Circle is an imaginary line that delimits the northern boundary of Antarctica. It is a distinctive parallel of latitude at approximately 66°30′ south. Thus it is located about 4590 mi (7345 km) south of the Equator and about 1630 mi…

Sea-level rise is caused by adding mass (water) and heat to the ocean. The Antarctic ice sheet represents a potentially large source of water. Knowing the speed and direction (velocity) at which the Antarctic ice sheet is flowing to the ocean is…

The Antarctic Ocean, sometimes called the Southern Ocean, is the watery belt surrounding Antarctica. It includes the great polar embayments of the Weddell Sea and Ross Sea, and the deep circumpolar belt of ocean between 50 and 60°S and the southern…

Any of several living mammals in the suborder Vermilingua in the order Pilosa that feed on ants and termites. The common name of anteater is assigned to four species of living mammals classified in the suborder Vermilingua in the order Pilosa…

The name given to a group of African and Asian hoofed ruminants of the order Artiodactyla. The 91 species in 31 genera are divided into five subfamilies (Bovinae, Cephalophinae, Hippotraginae, Antilopinae, and Caprinae) in the family Bovidae. The…

A magnesium-rich orthorhombic amphibole with perfect {210} cleavage and a color which varies from white to various shades of green and brown. It is a comparatively rare metamorphic mineral which occurs as slender prismatic needles, in fibrous…

A class of phylum Cnidaria (formerly known as Coelenterata) whose members are marine, live exclusively as polyps, and occur as solitary individuals or in clonal or colonial groups. Most adult anthozoans are attached to a firm object or the sea…

An order of Paleozoic tetrapods that arose Carboniferous (Mississippian) (345 million years ago) and lasted until the Early Triassic (about 245 MYA). The term anthracosaurs (meaning “coal reptiles”) encompasses a range of genera that may not all be…

A fold in layered rocks in which the strata are inclined down and away from the axes. The simplest anticlines (see illustration) are symmetrical, but in more highly deformed regions they may be asymmetrical, overturned, or recumbent. Most anticlines…

An order of cnidarians (subclass Hexacorallia) commonly known as black, or thorny, corals. Antipatharians have been identified in all of the world's oceans at depths of 4 to 8600 m (13 to 28,200 ft), but most known species live between 100 and 1000…

One of the three living orders of the class Amphibia, encompassing the frogs and toads. Because members of the order Anura (anurans) lack a tail, they differ most obviously from salamanders (order Urodela; also called Caudata). Usually, frogs and…

Apatite is a generic name used to describe calcium phosphates with the simplified formula Ca10(PO4)6X2 (X = F, Cl, OH). Three end members form a complete solid-solution series involving the halide and hydroxyl anions, and these are the species…

Any of the vermiform (wormlike) marine mollusks covered by a cuticle invested with innumerable calcareous scales or spines (sclerites) forming a scleritome. There are two groups that together are known as the Aplacophora: (1) Chaetodermomorpha or…

A fine-grained, sugary-textured rock, generally of granitic composition; also any body composed of such rock. GraniteThe light-colored rock consists chiefly of quartz, microcline, or orthoclase perthite and sodic plagioclase, with small amounts of…

The smallest order of the class Amphibia, consisting of the wormlike caecilians. The order Apoda is sometimes called Gymnophiona. Its members are known commonly as caecilians, which are wormlike, legless amphibians with indistinct or even hidden…

An order of birds consisting of two dissimilar groups, the swifts (Apodi) and the hummingbirds (Trochili). These birds have been placed together because of several anatomical specializations of the wings and feathers, although an alternative…

A hydrous calcium potassium silicate containing fluorine. The composition is variable but approximates to KFCa4(Si2O5)4· 8H2O. It resembles the zeolites, with which it is sometimes classified, but differs from most zeolites in having no aluminum. It…

A little-studied group of ciliates comprising an order of the Holotrichia. The majority occur as commensals on marine crustaceans. This association is generally more intimate and complex than that of the chonotrichs found on related hosts. Their…

A class of marine, planktonic animals in the subphylum Tunicata. The class Appendicularia (alternatively known as Larvacea) is characterized by the persistence of a tail, notochord, gill slits, and dorsal nerve cord throughout life and by a unique…

A subsurface zone that yields economically important amounts of water to wells. The term is synonymous with water-bearing formation. An aquifer may be porous rock, unconsolidated gravel, fractured rock, or cavernous limestone. Economically important…

The largest class of the subphylum Chelicerata in the phylum Arthropoda. Most familiar of the included orders in the class Arachnida are the spiders (Fig. 1), scorpions (Fig. 2), harvestmen, and mites and ticks. Members of the class Arachnida…

An order of nematodes in which the amphids are simple spirals that appear as elongate loops, shepherd's crooks, question marks, or circular forms. The cephalic sensilla are often separated into three circlets: the first two are papilliform or the…

An order of Paleozoic diapsid reptiles including the families Petrolacosauridae and Araeoscelidae. Members of these families resemble primitive modern lizards, such as the green iguana, in size and most body proportions, but are distinguished by…

One of three naturally occurring mineral forms of calcium carbonate (CaCO3). The other forms (or polymorphs) are the abundant mineral calcite and the relatively rare mineral vaterite. Still other forms of calcium carbonate are known, but only as…

A natural order of the class Arachnida, commonly known as the spiders. Members of the order Araneae (also called Araneida)—the spiders—are widespread over most of the land areas of the world and are well adapted to many different habitats. These…

An order of regular sea urchins (class Echinoidea) that have a stirodont Aristotle's lantern (jaw apparatus) with the epiphyses not meeting at the top of the lantern, and with keeled teeth that are T-shaped in cross section. The ambulacral plates of…

An order of Lobosia. The shell (test) of these protozoa has a single, well-defined aperture through which slender fingerlike pseudopodia (lobopodia) can be extended. The test often has an internal chitinous layer known as a glycocalyx and an…

An extinct group of mainly Lower Cambrian marine sponges which, although lacking spicules, possessed an intricate, highly porous skeleton of calcite. It was probably a monophyletic group; that is, all representatives were derived from a single…

An order of ancestrally wingless insects, commonly known as bristletails, in the subclass Apterygota; about 350 species are known in the two families, Machilidae and Meinertellidae. All have slender, spindle-shaped bodies of about 0.4–0.8 in. (1–2…

An order of extinct, free-sporing plants (cryptogams) that lived during the Late Devonian through Early Mississippian. The order includes the genera Archaeopteris (which has been extensively investigated), Actinoxylon, Actinopodium, Svalbardia,…

A genus of fossil birds in the extinct order Archaeopterygiformes, characterized by flight feathers like those of modern birds. Archaeopteryx is generally accepted as the oldest known fossil bird. It represents a unique snapshot of evolution; most…

The most diverse group of early Tertiary mammals are the archaic ungulates (condylarths). Although closely related, they are not a monophyletic group but, along with various extinct and extant mammals, form the taxon Ungulata (Fig. 1). They are…

A period of geologic time from about 4000 to 2500 million years ago (MYA). During the Archean Eon (Fig. 1) a large percentage of the Earth's continental crust formed, plate tectonics began, very warm climates and oceans existed, and life appeared on…

The establishment of the temporal sequence of prehistoric events. An event can be defined as a duration of time during which certain features came together to form a set, for example, when certain materials were formed into an object (manufacturing…

A class of the phylum Acanthocephala. The adult worms are parasitic in terrestrial vertebrates. The body wall and lemnisci of species in this class have numerous amitotically fragmented nuclei or a few ameboid giant nuclei. The main trunks of the…

A diversified group of diapsid reptiles, comprising dinosaurs (and their descendants, birds), crocodylians, pterosaurs (flying reptiles), and various forms closely related to these groups. In the past, the primitive archosaurs from the Triassic…

Defined primarily by climatic rather than latitudinal criteria, arctic islands are those in the Northern Hemisphere where the mean temperature of the warmest month does not exceed 50°F (10°C) and that of the coldest is not above 32°F (0°C).…

The parallel of latitude approximately 66°30′ north of the Equator, or 23.5° from the North Pole. Named for the northern constellation Bear, the Arctic Circle has the same angular distance from the Equator as the inclination of the Earth's axis from…

The Arctic Ocean is an enclosed ocean, connected to the Pacific Ocean through Bering Strait and Bering Sea, and to the Atlantic Ocean through Fram Strait and the Barents, Greenland, and Norwegian seas (see illustration). The Arctic Ocean is the…

The arenaceous rocks (arenites) include all those clastic sedimentary rocks whose particle sizes range from 2 to 0.06 mm, or if silt is included, to 0.004 mm. Some arenites are composed primarily of carbonate particles, in which case they are called…

An order of teleost fishes, comprising the argentines or herring smelts, in the superorder Protacanthopterygii. Members of the order Argentiniformes are marine, deep-water fishes. They are identifiable by the following combination of characters: a…

Clastic sedimentary rocks containing silt- or clay-sized particles that are less than 0.0625 mm and/or clay minerals. The argillaceous rocks (lutites) include shales, argillites, siltstones, and mudstones. They are the most abundant sedimentary rock…

A suborder of leeches (order Hirudinida) characterized by the absence of a protrusible proboscis and including both blood-feeding and macrophagous (predatory) varieties. Most notorious among the blood-feeding leeches in the Arhynchobdellida is the…

A heavily armored mammal in the order Cingulata. The order Cingulata (formerly Edentata or Xenarthra) comprises the armadillos, which are divided into two extant families: Chlamyphoridae and Dasypodidae. These creatures are characterized by bony…

Groundwater conditions formed by water-bearing rocks (aquifers) in which the water is confined above and below by impermeable beds. These systems are named after the province of Artois in France, where artesian wells were first observed.Because the…

The largest phylum in the animal kingdom, including the well-known insects, spiders, ticks, and crustaceans, as well as many smaller, less well-known groups. Members of the phylum Arthropoda, that is, arthropods, are the dominant metazoans…

The only surviving subclass of the echinoderm class Crinoidea. The Articulata include all living members of the echinoderm class Crinoidea and are generally regarded as the post-Paleozoic crinoids. Crinoids are exclusively marine organisms, as are…

Substitutes for sugar (sucrose) in foods and beverages that provide intense sweetness with few to no calories. Artificial sweeteners are sometimes called low-calorie sweeteners, high-intensity sweeteners, noncaloric sweeteners, non-nutritive…

An order comprising the even-toed ungulates (hoofed mammals). There are two main radiations: the predominantly omnivorous Bunodontia, including suoids (such as pigs, peccaries, and hippos); and the more herbivorous Selenodontia, including camels and…

A group of highly fibrous, naturally occurring minerals with separable, long and thin fibers. Asbestos fibers are strong and flexible enough to be spun and woven as well as resistant to heat and chemicals. Historically, asbestos was used for fire…

An order of nematodes in which the oral opening is generally surrounded by three or six labia; in some taxa, labia are absent, but the cephalic sensilla are always evident. Usually there are eight cephalic or labial sensilla; the submedians may be…

A class of Tunicata (Urochordata) which occurs as solitary zooids or, by a process of asexual budding, develops into colonies. Zooids of the class Ascidiacea vary in length from a few millimeters to 10 in. (25 cm). Individuals or colonies are…

A subtaxon of marine crustaceans in the subclass Thecostraca. Members of the Ascothoracica are ectoparasites or endoparasites of cnidarians and echinoderms. The body is enclosed in a voluminous saclike mantle that is up to 20 mm (0.8 in.) in length;…

The largest of the world's continents. With its peninsular extension, commonly called the continent of Europe, it is the major portion of the broad east-west extent of the Northern Hemisphere landmasses. In many ways Asia is more a cultural concept…

A white, crystalline compound, 1-aspartyl-1-phenylalanine methyl ester (APM), with formula (1). It is slightly soluble in water. Its sweetening properties were discovered accidentally in 1965 when the compound, a dipeptide, was produced as an…

A small group of trematodes (flukes) with about 80 freshwater and marine species in 4 families and 13 genera, infecting fishes (chondrichthyans and teleosts) and turtles. The Aspidogastrea vary in length from about 2 mm (0.08 in.) to several…

A small order of specialized holostean fishes which are first recorded from Middle Jurassic deposits of Europe. Later records indicate that they were a very successful group in terms of geographic range, and probably had a worldwide distribution in…

A layer of variable thickness within the upper mantle of the Earth that is mechanically weaker (less rigid) than both the overlying lithosphere and the underlying mesosphere. The depth of transition from lithosphere to asthenosphere can be as little…

An order of protozoans, subclass Holotrichia, in which all species are mouthless. All species are parasitic in other animals, typically oligochaete annelids. Many astomatids possess an elaborate holdfast organelle, of value in attachment to the…

A relatively small order of extinct South American ungulates in the infraclass Eutheria, known from the late Paleocene to the late Miocene. Members of the order Astrapotheria are customarily divided into two suborders: the late Paleocene–Eocene…

An order of actinopterygian (ray-finned) fishes consisting of only one family, Ateleopodidae, known as jellynose fishes. Ateleopodiformes along with the order Stomiiformes constitute a teleost superorder called the Stenopterygii. Jellynose fishes…

An order of Granuloreticulosia in which the naked amebas form branched, threadlike, interconnected pseudopodia (reticulopodia). Species are known from freshwater, salt water, and brackish water. General characteristics of the order Athalamida are…

An order of actinopterygian (ray-finned) fishes, including the silversides, rainbowfishes, and blue eyes. The members of the Atheriniformes, together with those of the Beloniformes and Cyprinodontiformes, comprise the series Atherinomorpha.…

The large body of seawater separating the continents of North and South America in the west from Europe and Africa in the east and extending south from the Arctic Ocean to the continent of Antarctica. The Atlantic is the second-largest ocean water…

A gaseous layer that envelops the Earth and most other planets in the solar system. Earth, Venus, Mars, Jupiter, Saturn, Uranus, Neptune, and Titan (Saturn's largest satellite) are all known to possess substantial atmospheres that are held by the…

A scientific discipline concerned with the chemical composition of the Earth's atmosphere. Topics falling within the field include the emission, transport, and deposition of atmospheric chemical species; the rates and mechanisms of chemical…

An annular coral reef, with or without small islets, that surrounds a lagoon without projecting land area. Physiography Most atolls are isolated reefs that rise from the deep sea. They vary considerably in size and shape. Small rings, some without…

A group of monoclinic calcic pyroxenes which have the general chemical formula (Ca,Mg,Fe)(Mg,Fe)-Si2O6, in which calcium is the dominant cation in the first cation position. Monoclinic pyroxene with substantial iron or magnesium in place of calcium…

An order of about 219 species of teleost fishes ranked in four suborders. The order Aulopiformes is distinguished by certain skeletal specializations of the gill arches that are unknown in other teleost fishes. They share characters with the order…