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A class of the phylum Radiozoa in the Actinopoda. The kingdom Protozoa contains 18 phyla. One of the parvkingdoms (a hierarchical classification between kingdom and superphylum that is controversial and not officially recognized as such) is the…

An order of ectoparasitic annelid worms consisting of one species, Acanthobdella peledina, which feeds on the dermal tissues of freshwater salmonid fishes in the far regions of Northern Europe. The order Acanthobdellida is taxonomically odd in that…

A distinct phylum of helminths (parasitic worms), commonly known as the spiny- or thorny-headed worms. The phylum Acanthocephala comprises three classes: Archiacanthocephala, Palaeacanthocephala, and Eoacanthocephala (a suggested fourth class,…

A taxon of marine protozoans belonging to the Acantharea. Acanthometrida (also known as Arthracanthida) is a taxonomic grouping (sometimes considered as an order or subclass) placed within the Acantharea (Acantharia) taxon. All members are…

A taxon of arachnids (traditionally considered a subclass of the Arachnida), comprising the mites and ticks. The Acari (also Acarina) consist of two superorders of small to minute arachnids. They have two body regions (Fig. 1)—a headlike gnathosoma…

An order of actinopterygian (ray-finned) fishes represented by the sturgeons and paddlefishes. The order Acipenseriformes, the only extant order of about 10–12 orders of chondrosteans (subclass Chondrostei), probably originated from palaeonisciform…

A group of small, soft-bodied, ciliated worms with the form (“morph”) of acoel turbellarians. The Acoelomorpha is now ranked as a phylum, distinct from other turbellarian worms of the phylum Platyhelminthes. Its relation to other animals, however,…

A subtaxon of burrowing barnacles in the subclass Thecostraca. All members of the Acrothoracica burrow into shells of mollusks and thoracican barnacles, echinoderm tests, bryozoans, dead coral, and limestone. The entrance to the burrow is slitlike.…

An order of inarticulated brachiopods. Together with Lingulida and Siphonotretida, Acrotretida is one of the three orders of inarticulated brachiopods currently classified in class Lingulata in subphylum Linguliformea (phylum Brachiopoda).…

An order of the class Anthozoa, commonly known as the sea anemones. Actiniaria contains approximately 1100 species. They are strictly marine, occurring from northern to southern polar latitudes and inhabiting regions that range from the intertidal…

An order of Heliozoia. These protozoans are characterized by stiff arms radiating from the central cell mass. Each arm is supported internally by an array of microtubules which terminate on the surface of nuclei. The organisms are found in…

A class of Sarcodina; or in some modern schemes a superclass (Actinopoda) within the phylum Sarcomastigophora. Some of these protozoans have more or less permanent pseudopodia, composed of a central shaft of microtubules surrounded by a thin…

A class of teleostom fishes commonly known as the ray-finned fishes, which are distinguished by the structure of the paired fins that are supported by dermal rays. The class Actinopterygii comprises the subclasses Cladistia [Polypteriformes (bichirs…

One of the four fundamental operations of arithmetic and algebra. The symbol + of addition is thought to be a ligature for “et,” used in a German manuscript of 1456 to denote addition. Its first printed appearance is in Johann Widman's Behennede und…

Aeronomy concentrates on the regions above the tropopause or upper part of the atmosphere. The region of the atmosphere below the tropopause is the site of most of the weather phenomena that so directly affect all life on the planet; this region has…

A variety of chalcedonic quartz that is distinguished by the presence of color banding in curved or irregular patterns (Fig. 1). Most agate used for ornamental purposes is composed of two or more tones or intensities of brownish-red, often…

A large, rabbit-sized rodent, belonging to the family Dasyproctidae and inhabiting South America and Central America, which is well adapted for speed because of its elongated hind legs. The agouti is a large rodent that resembles the rabbit or hare…

A sodium-rich plagioclase feldspar mineral whose composition extends over the range Ab100An0 to Ab90An10, where Ab (= albite) is NaAlSi3O8 and An (= anorthite) is CaAl2Si2O8 (see illustration). Albite occurs in crustal igneous rocks as a major…

An order of actinopterygian fishes in the subdivision Elopomorpha, along with Elopiformes, Anguilliformes, and Saccopharyngiformes, all of which have a leptocephalous larval stage. Albuliformes, comprising three families, eight genera, and 30…

An order of the class Anthozoa (phylum Cnidaria), containing 2500–3000 species, commonly referred to as soft corals, sea fans, sea whips, and precious coral. Members of the Alcyonacea are strictly marine. They occur from northern to southern polar…

The branch of mathematics dealing with the solution of equations. These equations involve unknowns, or variables, along with fixed numbers from a specified system (Fig. 1). The origins of algebra were based on the need to develop equations that…

The study of zero sets of polynomial equations. Examples are the parabola y − x2 = 0, thought of as sitting in the (x, y)-plane, and the locus of all points (t, t2, t3, t4), which is defined by Eqs. (1), 1 in the coordinate space with coordinates w,…

Light emissions in the upper atmosphere from elemental lithium, potassium, and especially sodium. These alkali metals are present in the upper atmosphere at altitudes from about 50 to 62 mi (80 to 100 km) and are very efficient in resonant…

A large aquatic reptile of the family Alligatoridae. Alligators belong to the family Alligatoridae in the reptilian order Crocodylia (Crocodilia). Common usage generally restricts the name alligator to the two extant species of the genus Alligator.…

One of the four subclasses of Mammalia, containing a single order, the Multituberculata. The Allotheria first appeared in the Late Jurassic and survived well into the Cenozoic, a period of at least 100,000,000 years. Fossils are known from North…

Vicugna pacos, a member of the camel family (Camelidae), which has been domesticated for more than 2000 years and is economically important for its long, fine wool. The alpaca (Vicugna pacos; formerly, Lama pacos) [see illustration] belongs to the…

A mineral of composition KAl3(SO4)2(OH)6. Alunite occurs in white to gray rhombohedral crystals or in fine-grained, compact masses. It has a hardness of 4 and a specific gravity of 2.6–2.8.Alunite is produced by sulfurous vapors on acid volcanic…

Most commonly, a generic name for all fossil resins, although it has been restricted by some to refer only to succinite, the mineralogical species of fossil resin that includes most of the Baltic Coast deposits. Resins generally are complex mixtures…

A fatty substance formed in the intestinal tract of the sperm whale (Physeter catodon). There is a question as to whether the origin of ambergris is normal or pathological, but it does serve as protection from the horny indigestible portions of the…

The tailless whip scorpions or whip spiders, an order of the class Arachnida. There are about 80 species in the tropics and subtropics. They are flattened, red to brown, and range from 5 to 45 mm (0.2 to 1.8 in.) in body length. All are nocturnal,…

Any protozoan moving by means of protoplasmic flow. In their entirety, the ameboid protozoa include naked amebas, those enclosed within a shell or test, as well as more highly developed representatives such as the heliozoians, radiolarians, and…

The transparent purple to violet variety of the mineral quartz. Although quartz is perhaps the commonest gem mineral known, amethyst is rare in the deep colors that characterize fine quality. Amethyst is usually colored unevenly and is often heated…

An order of actinopterygian fishes in the subclass Neopterygii. Amiiformes plus several related fossil orders comprise the Halecomorphi, a well-developed group known from the middle Mesozoic. The Amiiformes comprise several families that persisted…

Determination of the relative or absolute age of materials or objects by measurement of the degree of racemization of the amino acids present. With the exception of glycine, the amino acids found in proteins can exist in two isomeric forms called d-…

A collective term for the classes Reptilia (reptiles), Aves (birds), and Mammalia (mammals) of the subphylum Vertebrata. The remaining vertebrates, including the several classes of fishes and the amphibians, are grouped together as the Anamnia.…

An order of Lobosia without protective coverings (tests). These protozoa range in size from about 4 micrometers to 0.08–0.12 in. (2–3 mm). Pellicles may be thin, as in Amoeba proteus, or thicker and less flexible, as in Thecamoeba verrucosa (Fig.…

One of four classes (the others are Reptilia, Aves, and Mammalia) composing the superclass Tetrapoda of the subphylum Vertebrata. The living amphibians number over 4700 species and are classified in three orders: Salientia or Anura (frogs and toads,…

Collectively, members of the class Amphibia are facing massive reductions in total numbers, and the overall biodiversity of amphibians across the globe is under threat. Scientists have determined that global climate change and habitat destruction…

A mineral with a framework structure in which all the aluminosilicate tetrahedral vertices are linked, thus allying it to the feldspars, feldspathoids, and zeolites. The structure is cubic, space group Ia3d, a = 1.371 mm. Its formula is…

Total variation in experimental data is partitioned into components assignable to specific sources by the analysis of variance. This statistical technique is applicable to data for which (1) effects of sources are additive, (2) uncontrolled or…

A branch of mathematics in which algebra is applied to the study of geometry. Because algebraic methods were first systematically applied to geometry in 1637 by the French philosopher-mathematician René Descartes, the subject is also called…

A framework for solving a problem. The analytic hierarchy process is a systematic procedure for representing the elements of any problem. It organizes the basic rationality by breaking down a problem into its smaller constituents and then calls for…

A collective term, also referred to as Anamniota, for the vertebrate animals that lack an amnion in development, including the Agnatha, Chondrichthyes, Osteichthyes, and Amphibia. The amnion is a protective embryonic envelope that encloses the…

Formerly, a group of reptiles that was recognized in rank-based classifications. Amniotes were once divided into subclasses, based on the patterns of temporal fenestrae (openings in the skull roof posterior to the openings for the eyes). Subclass…

An extant order of the crustacean superorder Syncarida confined to freshwater and recorded from both surface and subterranean waters. Members of the order Anaspidacea (anaspidaceans) lack a carapace. The first thoracomere (thoracic somite) is…

A nesosilicate mineral, composition Al2SiO5, crystallizing in the orthorhombic system. It occurs commonly in large, nearly square prismatic crystals. The variety chiastolite has inclusions of dark-colored carbonaceous material arranged in a regular…

A plagioclase feldspar with composition Ab70An30 to Ab50An50 (Ab = NaAlSi3O8; An = CaAl2Si2O8). Andesine occurs primarily in igneous rocks, often in a glassy matrix as small, chemically zoned, lathlike crystals known as microlites. The rock types…

A typical volcanic rock erupted from a volcano associated with convergent plate boundaries. The process of subduction, which defines convergent plate boundaries, results in the sinking of oceanic lithosphere beneath either oceanic lithosphere or…

A mineral with the chemical composition PbSO4. Anglesite occurs in white or gray, orthorhombic, tabular or prismatic crystals or compact masses (see illustration). It is a common secondary mineral, usually formed by the oxidation of galena. Fracture…

The true eels, a large order of actinopterygian fishes, also known as the Apodes. The order Anguilliformes is related to Saccopharyngiformes (sackpharynx fishes), Elopiformes (tarpons), and Notacanthiformes (spiny eels and halosaurs); all members…

A mineral with the chemical composition CaSO4. Anhydrite occurs commonly in white and grayish granular masses, rarely in large, orthorhombic crystals (see illustration). Fracture is uneven and luster is pearly to vitreous. Hardness is 3–3.5 on Mohs…

An animal's movement through the atmosphere sustained by aerodynamic reaction or other forces. Animal flight includes gliding and flapping flight. Gliding flight does not involve the generation of thrust, whereas flapping flight does generate thrust…

The classification rank comprising the animals. Animals are taxonomically assigned to the classification kingdom known as Animalia. (The other kingdoms for classification of organisms include Plantae, Fungi, Protista, and Monera; in certain…

The carbonate mineral Ca(Fe,Mg)(CO3)2, also commonly containing some manganese. The mineral has hexagonal (rhombohedral) symmetry (see illustration) and has the cation-ordered structure of dolomite. The name is applied only to those species in which…

The phylum comprising the multisegmented, invertebrate wormlike animals, of which the most numerous are the marine bristle worms and the most familiar the terrestrial earthworms. The Annelida (meaning little annuli or rings) include the Polychaeta…

The calcium-rich plagioclase feldspar with composition Ab0An100 to Ab10An90 (Ab = NaAlSi3O8; An = CaAl2Si2O8), occurring in olivine-rich igneous rocks and rare volcanic ejecta (for example, at Mount Vesuvius, Italy, and Miyakejima, Japan). Hardness…

The name usually given to alkali feldspars which have a chemical composition ranging from Or40Ab60 to Or10Ab90± up to approximately 20 mole % An (Or, Ab, An = KAlSi3O8, NaAlSi3O8, CaAl2Si2O8) and which deviate in one way or another from monoclinic…

A rock composed of 90 vol % or more of plagioclase feldspar. Strictly, the rock is composed entirely of crystals discernible with the eye, but some finely crystalline examples from the Moon have been called anorthosite or anorthositic breccia. Two…

An order of branchiopod crustaceans, known as fairy shrimps and, in some cases, brine shrimps (Artemia). Members of the order Anostraca (anostracans) range up to about 100 mm (4 in.) in length but usually are much smaller. The trunk consists of…

Oxygen-depleted regions in marine environments. The dynamic steady state between oxygen supply and consumption determines the oxygen concentration. In regions where the rate of consumption equals or exceeds the rate of supply, seawater becomes…

An order of birds comprising two families, the screamers (Anhimidae) of South America and the worldwide waterfowl (Anatidae). They are closely related to the Galliformes (fowls), with the screamers being a rather intermediate group. The waterfowl…

The Antarctic Circle is an imaginary line that delimits the northern boundary of Antarctica. It is a distinctive parallel of latitude at approximately 66°30′ south. Thus it is located about 4590 mi (7345 km) south of the Equator and about 1630 mi…

Sea-level rise is caused by adding mass (water) and heat to the ocean. The Antarctic ice sheet represents a potentially large source of water. Knowing the speed and direction (velocity) at which the Antarctic ice sheet is flowing to the ocean is…

The Antarctic Ocean, sometimes called the Southern Ocean, is the watery belt surrounding Antarctica. It includes the great polar embayments of the Weddell Sea and Ross Sea, and the deep circumpolar belt of ocean between 50 and 60°S and the southern…

Any of several living mammals in the suborder Vermilingua in the order Pilosa that feed on ants and termites. The common name of anteater is assigned to four species of living mammals classified in the suborder Vermilingua in the order Pilosa…

The name given to a group of African and Asian hoofed ruminants of the order Artiodactyla. The 91 species in 31 genera are divided into five subfamilies (Bovinae, Cephalophinae, Hippotraginae, Antilopinae, and Caprinae) in the family Bovidae. The…

A magnesium-rich orthorhombic amphibole with perfect {210} cleavage and a color which varies from white to various shades of green and brown. It is a comparatively rare metamorphic mineral which occurs as slender prismatic needles, in fibrous…

A class of phylum Cnidaria (formerly known as Coelenterata) whose members are marine, live exclusively as polyps, and occur as solitary individuals or in clonal or colonial groups. Most adult anthozoans are attached to a firm object or the sea…

A fold in layered rocks in which the strata are inclined down and away from the axes. The simplest anticlines (see illustration) are symmetrical, but in more highly deformed regions they may be asymmetrical, overturned, or recumbent. Most anticlines…

An order of cnidarians (subclass Hexacorallia) commonly known as black, or thorny, corals. Antipatharians have been identified in all of the world's oceans at depths of 4 to 8600 m (13 to 28,200 ft), but most known species live between 100 and 1000…

One of the three living orders of the class Amphibia, encompassing the frogs and toads. Because members of the order Anura (anurans) lack a tail, they differ most obviously from salamanders (order Urodela; also called Caudata). Usually, frogs and…

Apatite is a generic name used to describe calcium phosphates with the simplified formula Ca10(PO4)6X2 (X = F, Cl, OH). Three end members form a complete solid-solution series involving the halide and hydroxyl anions, and these are the species…

Any of the vermiform (wormlike) marine mollusks covered by a cuticle invested with innumerable calcareous scales or spines (sclerites) forming a scleritome. There are two groups that together are known as the Aplacophora: (1) Chaetodermomorpha or…

A fine-grained, sugary-textured rock, generally of granitic composition; also any body composed of such rock. GraniteThe light-colored rock consists chiefly of quartz, microcline, or orthoclase perthite and sodic plagioclase, with small amounts of…

The smallest order of the class Amphibia, consisting of the wormlike caecilians. The order Apoda is sometimes called Gymnophiona. Its members are known commonly as caecilians, which are wormlike, legless amphibians with indistinct or even hidden…

An order of birds consisting of two dissimilar groups, the swifts (Apodi) and the hummingbirds (Trochili). These birds have been placed together because of several anatomical specializations of the wings and feathers, although an alternative…

A hydrous calcium potassium silicate containing fluorine. The composition is variable but approximates to KFCa4(Si2O5)4· 8H2O. It resembles the zeolites, with which it is sometimes classified, but differs from most zeolites in having no aluminum. It…

A little-studied group of ciliates comprising an order of the Holotrichia. The majority occur as commensals on marine crustaceans. This association is generally more intimate and complex than that of the chonotrichs found on related hosts. Their…

A class of marine, planktonic animals in the subphylum Tunicata. The class Appendicularia (alternatively known as Larvacea) is characterized by the persistence of a tail, notochord, gill slits, and dorsal nerve cord throughout life and by a unique…

A subsurface zone that yields economically important amounts of water to wells. The term is synonymous with water-bearing formation. An aquifer may be porous rock, unconsolidated gravel, fractured rock, or cavernous limestone. Economically important…

The largest class of the subphylum Chelicerata in the phylum Arthropoda. Most familiar of the included orders in the class Arachnida are the spiders (Fig. 1), scorpions (Fig. 2), harvestmen, and mites and ticks. Members of the class Arachnida…

An order of nematodes in which the amphids are simple spirals that appear as elongate loops, shepherd's crooks, question marks, or circular forms. The cephalic sensilla are often separated into three circlets: the first two are papilliform or the…

One of three naturally occurring mineral forms of calcium carbonate (CaCO3). The other forms (or polymorphs) are the abundant mineral calcite and the relatively rare mineral vaterite. Still other forms of calcium carbonate are known, but only as…

A natural order of the class Arachnida, commonly known as the spiders. Members of the order Araneae (also called Araneida)—the spiders—are widespread over most of the land areas of the world and are well adapted to many different habitats. These…

An order of regular sea urchins (class Echinoidea) that have a stirodont Aristotle's lantern (jaw apparatus) with the epiphyses not meeting at the top of the lantern, and with keeled teeth that are T-shaped in cross section. The ambulacral plates of…

An order of Lobosia. The shell (test) of these protozoa has a single, well-defined aperture through which slender fingerlike pseudopodia (lobopodia) can be extended. The test often has an internal chitinous layer known as a glycocalyx and an…

An extinct group of mainly Lower Cambrian marine sponges which, although lacking spicules, possessed an intricate, highly porous skeleton of calcite. It was probably a monophyletic group; that is, all representatives were derived from a single…

An order of ancestrally wingless insects, commonly known as bristletails, in the subclass Apterygota; about 350 species are known in the two families, Machilidae and Meinertellidae. All have slender, spindle-shaped bodies of about 0.4–0.8 in. (1–2…

A period of geologic time from about 4000 to 2500 million years ago (MYA). During the Archean Eon (Fig. 1) a large percentage of the Earth's continental crust formed, plate tectonics began, very warm climates and oceans existed, and life appeared on…

A class of the phylum Acanthocephala. The adult worms are parasitic in terrestrial vertebrates. The body wall and lemnisci of species in this class have numerous amitotically fragmented nuclei or a few ameboid giant nuclei. The main trunks of the…

A diversified group of diapsid reptiles, comprising dinosaurs (and their descendants, birds), crocodylians, pterosaurs (flying reptiles), and various forms closely related to these groups. In the past, the primitive archosaurs from the Triassic…

Defined primarily by climatic rather than latitudinal criteria, arctic islands are those in the Northern Hemisphere where the mean temperature of the warmest month does not exceed 50°F (10°C) and that of the coldest is not above 32°F (0°C).…

The parallel of latitude approximately 66°30′ north of the Equator, or 23.5° from the North Pole. Named for the northern constellation Bear, the Arctic Circle has the same angular distance from the Equator as the inclination of the Earth's axis from…

The Arctic Ocean is an enclosed ocean, connected to the Pacific Ocean through Bering Strait and Bering Sea, and to the Atlantic Ocean through Fram Strait and the Barents, Greenland, and Norwegian seas (see illustration). The Arctic Ocean is the…

The superficial contents of a geometrical figure of two dimensions. The area of any rectangle or square is the product of two adjacent sides, one of which may be called the base and the other the altitude. In general, any line segment that partially…

The arenaceous rocks (arenites) include all those clastic sedimentary rocks whose particle sizes range from 2 to 0.06 mm, or if silt is included, to 0.004 mm. Some arenites are composed primarily of carbonate particles, in which case they are called…

An order of teleost fishes, comprising the argentines or herring smelts, in the superorder Protacanthopterygii. Members of the order Argentiniformes are marine, deep-water fishes. They are identifiable by the following combination of characters: a…

Clastic sedimentary rocks containing silt- or clay-sized particles that are less than 0.0625 mm and/or clay minerals. The argillaceous rocks (lutites) include shales, argillites, siltstones, and mudstones. They are the most abundant sedimentary rock…

A suborder of leeches (order Hirudinida) characterized by the absence of a protrusible proboscis and including both blood-feeding and macrophagous (predatory) varieties. Most notorious among the blood-feeding leeches in the Arhynchobdellida is the…

A branch of mathematics dealing with numbers, operations on numbers, and computation. Arithmetic is useful in solving many practical problems, such as buying, selling, budgets, sports statistics, and measurement. The usual numbers of arithmetic are…