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A plant, Musa textilis, of the banana family that is valuable for its hard fiber (also known as Manila hemp) and native to Borneo and the Philippines. Abaca is one of the strongest of the hard fibers. Commercially, it is known as Manila hemp. The…

A family of adenosine triphosphate–binding cassette (ABC) transmembrane proteins that use energy to transport various molecules across extracellular and intracellular membranes (cytoplasmic membranes, endoplasmic reticulum, mitochondria, or…

One of the major plant hormones that plays an important role in plant growth and development. Abscisic acid (ABA) is ubiquitous in vascular plants; the highest levels are found in young leaves and in developing fruits and seeds. Abscisic acid is…

The net movement (transport) of water and solutes from outside an organism to its interior. The unidirectional flow of materials into an animal from the environment generally takes place across the alimentary tract, the lungs, or the skin, and in…

A naturally occurring quaternary ammonium cation ester, with the formula CH3(O)COC2H4N(CH3)3+, which plays a prominent role in nervous system function as a neurotransmitter. Acetylcholine (Fig. 1) is an organic compound released from certain…

An order of monocotyledonous angiosperms composed of a single genus, Acorus (sweet flag or sweet calamus), with two species widespread in the Northern Hemisphere. Formerly, Acorus was included in the aroid family, Araceae, but several lines of…

A viral disease of humans caused by the human immunodeficiency virus (HIV), which attacks and compromises the body's immune system. Individuals infected with the human immunodeficiency virus (HIV) proceed through a spectrum of stages that ultimately…

An induced state in which antigens originally regarded as foreign become regarded as self by the immune system. Tolerance can be induced (tolerization) in all of the cells of the immune system, including T cells (also known as T lymphocytes), the…

A genus of gram-negative, immotile and nonspore-forming, oval to rod-shaped, often pleomorphic bacteria which occur as parasites or pathogens in mammals (including humans), birds, and reptiles. They are facultatively aerobic, capable of fermenting…

A heterogeneous collection of bacteria with diverse ecologies and physiologies. The initial definition was based on their branching filamentous cellular morphology during a stage of the growth cycle. The name actinomycete means “ray fungus,” and for…

A characteristic of an organism that makes it fit for its environment or for its particular way of life. Adaptation is a key biological process in organisms and can take many forms. Typically, adaptation refers to two mechanisms: (1) an adjustment…

The science of optical systems in which a controllable optical element, usually a deformable mirror, is used to optimize the performance of the system, for example, to maintain a sharply focused image in the presence of wavefront aberrations. A…

A type of defective virus particle in which part of the genetic material of papovavirus SV40 is encased within an adenovirus protein coat (capsid). Human adenoviruses require human cells for their propagation; however, papovavirus SV40 can serve as…

A family of double-stranded DNA viruses, collectively known as adenoviruses, which are icosahedrally symmetric, usually found in the respiratory tract of the host species, and often associated with respiratory diseases. Members of the Adenoviridae…

A complex endocrine organ in proximity to the kidney. Adrenal gland tissue is present in all vertebrates from cyclostomes to placental mammals. The adrenal consists of two functionally distinct tissues: steroidogenic cells of mesodermal origin and…

A bacterial genus in the family Vibrionaceae comprising oxidase-positive, facultatively anaerobic, monotrichously flagellated gram-negative rods. The mesophilic species are A. hydrophila, A. caviae, and A. sobria; the psychrophilic one is A.…

Aeronomy concentrates on the regions above the tropopause or upper part of the atmosphere. The region of the atmosphere below the tropopause is the site of most of the weather phenomena that so directly affect all life on the planet; this region has…

A variety of chalcedonic quartz that is distinguished by the presence of color banding in curved or irregular patterns (Fig. 1). Most agate used for ornamental purposes is composed of two or more tones or intensities of brownish-red, often…

A reaction in which suspended particles are aggregated or clumped. It occurs upon the admixture of another type of particle, a change in the composition of the suspending fluid, or the addition of a soluble agent that acts as a bridge between two or…

A substance that will cause a clumping of particles such as bacteria or erythrocytes. Of major importance are the specific or immune agglutinins, which are antibodies that will agglutinate bacteria containing the corresponding antigens on their…

An age-contingent reduction in physiological function and activity, with a concomitant increase in mortality rate and a diminution in reproductive rate. Aging is the process of becoming older, which is influenced by genetic and environmental…

Aircraft specifically designed or adapted from a general utility airframe for use in agriculture and forestry and for control of insect vectors of human, animal, and plant diseases. Agricultural aircraft (Fig. 1) have become an indispensable tool…

A discipline concerned with solving the engineering problems of providing food and fiber for the people of the world. These problems include designing agricultural machinery to work the soil and harvest the crops, as well as developing water…

The science that deals with the selection, breeding, nutrition, and management of domestic animals for economical production of meat, milk, eggs, wool, hides, and other animal products. The broad field of agricultural science focuses on two realms:…

The techniques and methods used in plowing, harrowing, planting, tilling, harvesting, drying, storing, and processing of agricultural crops. Agricultural soil and crop practices seek to keep the quality and fertility of soils at high levels so as to…

A model for the functioning of an agricultural system, with all inputs and outputs. An ecosystem may be as small as a set of microbial interactions that take place on the surface of roots, or as large as the globe. An agroecosystem may be at the…

The science and study of crops and soils. Agronomy is the umbrella term for a number of technical research and teaching activities (see illustration): crop physiology and management, soil science, plant breeding, and weed management frequently are…

The efficiency of a surface in reflecting light. White surfaces have albedos close to 1; black surfaces have albedos close to 0. Because this efficiency can vary with the direction to the light source, the direction to the observer, and the color…

A sodium-rich plagioclase feldspar mineral whose composition extends over the range Ab100An0 to Ab90An10, where Ab (= albite) is NaAlSi3O8 and An (= anorthite) is CaAl2Si2O8 (see illustration). Albite occurs in crustal igneous rocks as a major…

A plasma protein produced by the liver that maintains fluid balance in the blood and transports fatty acids in the plasma and interstitial fluid. It is the most abundant protein in human serum, and one of the first discovered and earliest studied…

Any of the deciduous shrubs and trees of the genus Alnus in the birch family (Betulaceae). There are approximately 30 species of alder, with 10 in the United States. Alders are widespread in cool north temperate regions and are also found southward…

The world's most valuable forage legume, Medicago sativa, also known as lucerne, and less often as purple medic, medica, snail clover, median herb, Burgundy hay or clover, and Chilean clover. Alfalfa is often referred to as the queen of forages. It…

Light emissions in the upper atmosphere from elemental lithium, potassium, and especially sodium. These alkali metals are present in the upper atmosphere at altitudes from about 50 to 62 mi (80 to 100 km) and are very efficient in resonant…

A fluid-filled sac- or sausagelike, extraembryonic membrane lying between the outer chorion and the inner amnion and yolk sac of the embryos of reptiles, birds, and mammals. The allantois eventually fills up the space of the extraembryonic coelom in…

Any of a number of alternative forms of a gene. Allele is a contraction of allelomorph, a term that William Bateson used to designate one of the alternative forms of a unit showing mendelian segregation. New alleles arise from existing ones by…

Altered reactivity in humans and other animals to allergens (substances foreign to the body that trigger an allergic response) induced by exposure through injection, inhalation, ingestion, or skin contact. An allergy (or allergic reaction; Fig. 1)…

The study of changes in the characteristics of organisms with body size. Characteristics such as body parts or timing of reproductive events do not necessarily change in direct proportion to body size, and the ways in which they change relative to…

The dried, unripe fruits (berries) of a small, tropical, evergreen tree, Pimenta dioica (P. officinalis), of the myrtle family (Myrtaceae). Allspice (see illustration), which is also known as Jamaica pepper, is a native of the West Indies and parts…

A book that contains astronomical or meteorological data arranged according to days, weeks, and months of a given year and may also include diverse information of a nonastronomical character. This article is restricted to astronomical and…

Prunus dulcis (also known as Prunus amygdalus or Amygdalus communis), a small deciduous tree of the order Rosales that is closely related to the peach and other stone fruit trees, and grown widely for its ellipsoidal edible seeds. The fruit of the…

Vicugna pacos, a member of the camel family (Camelidae), which has been domesticated for more than 2000 years and is economically important for its long, fine wool. The alpaca (Vicugna pacos; formerly, Lama pacos) [see illustration] belongs to the…

A glycoprotein that is normally present in significant amounts only in the serum of the fetus. It is produced in the yolk sac, the liver, and other tissues of the gastrointestinal tract. Its role is unknown, but alpha fetoprotein may function as a…

A mineral of composition KAl3(SO4)2(OH)6. Alunite occurs in white to gray rhombohedral crystals or in fine-grained, compact masses. It has a hardness of 4 and a specific gravity of 2.6–2.8.Alunite is produced by sulfurous vapors on acid volcanic…

An annual plant (seldom perennial) of the genus Amaranthus (family Amaranthaceae) that is distributed worldwide in warm and humid regions. Amaranth plants are botanically distinguished by their small chaffy flowers (see illustration), which are…

Most commonly, a generic name for all fossil resins, although it has been restricted by some to refer only to succinite, the mineralogical species of fossil resin that includes most of the Baltic Coast deposits. Resins generally are complex mixtures…

The transparent purple to violet variety of the mineral quartz. Although quartz is perhaps the commonest gem mineral known, amethyst is rare in the deep colors that characterize fine quality. Amethyst is usually colored unevenly and is often heated…

Any of the organic compounds possessing one or more basic amino groups and one or more acidic carboxyl groups. Of the more than 80 amino acids that have been found in living organisms, approximately 20 serve as the building blocks for proteins.…

Determination of the relative or absolute age of materials or objects by measurement of the degree of racemization of the amino acids present. With the exception of glycine, the amino acids found in proteins can exist in two isomeric forms called d-…

A thin, cellular, extraembryonic membrane forming a closed sac which surrounds the embryo in all reptiles, birds, and mammals and is present only in these forms; hence the collective term amniotes is applied to these animals. The amnion contains a…

A collective term for the classes Reptilia (reptiles), Aves (birds), and Mammalia (mammals) of the subphylum Vertebrata. The remaining vertebrates, including the several classes of fishes and the amphibians, are grouped together as the Anamnia.…

An enzyme that is essential for maintenance of cellular energy balance. All living cells carry out many energy-requiring processes, including growth and division, movement, and uptake of nutrients from their environment. To achieve this, they…

A stimulating drug that affects the brain and the body. Amphetamine is a stimulant of the central nervous system. Chemically, amphetamine is a racemic mixture of the l and d isomers of α-methyl-β-phenethylamine (see illustration). The l isomer has…

A disorder characterized by the accumulation of an unusual extracellular fibrous protein (amyloid) in the connective tissue of the body. The deposition of amyloid may be widespread, involving major organs and leading to serious clinical…

A mineral with a framework structure in which all the aluminosilicate tetrahedral vertices are linked, thus allying it to the feldspars, feldspathoids, and zeolites. The structure is cubic, space group Ia3d, a = 1.371 mm. Its formula is…

A collective term, also referred to as Anamniota, for the vertebrate animals that lack an amnion in development, including the Agnatha, Chondrichthyes, Osteichthyes, and Amphibia. The amnion is a protective embryonic envelope that encloses the…

A generalized or localized tissue reaction occurring within minutes of an antigen-antibody reaction. Similar reactions elicited by nonimmunologic mechanisms are termed anaphylactoid reactions. The term anaphylaxis was introduced by C. R. Richet to…

A nesosilicate mineral, composition Al2SiO5, crystallizing in the orthorhombic system. It occurs commonly in large, nearly square prismatic crystals. The variety chiastolite has inclusions of dark-colored carbonaceous material arranged in a regular…

A plagioclase feldspar with composition Ab70An30 to Ab50An50 (Ab = NaAlSi3O8; An = CaAl2Si2O8). Andesine occurs primarily in igneous rocks, often in a glassy matrix as small, chemically zoned, lathlike crystals known as microlites. The rock types…

A typical volcanic rock erupted from a volcano associated with convergent plate boundaries. The process of subduction, which defines convergent plate boundaries, results in the sinking of oceanic lithosphere beneath either oceanic lithosphere or…

Synthetic manufactured drugs derived from the male sex hormone testosterone. Androgenic-anabolic steroids, often termed simply as anabolic steroids, are artificially produced testosterone variants. They can provide a number of medical benefits when…

A class of steroid hormones produced in the testes and adrenal cortex that act to regulate masculine secondary sexual characteristics. The major naturally occurring androgens, such as testosterone, are C-19 steroids (that is, they each contain 19…

A prominently located constellation in the northern sky (see illustration), named for the daughter of Cassiopeia in Greek mythology: When Cassiopeia bragged that her daughter Andromeda was more beautiful than Poseidon's daughters, the Nereids,…

The giant spiral galaxy closest to Earth. The Andromeda Galaxy is seen in the constellation of Andromeda, which is visible from Earth in the autumn skies. The galaxy is also known as M31 and NGC 224, based on its entries in two widely used catalogs.…

A medical condition resulting from a reduction in the total quantity of hemoglobin or of red blood cells in the circulation. Anemia is a disorder marked by significant decreases in hemoglobin concentration and in the number of circulating red blood…

The condition of exhibiting no immune response to an antigen. Anergy is a self-tolerance mechanism in which the immune system shows unresponsiveness following an encounter with receptor-recognized antigens. Typically, lymphocytes (the B and T cells…

The origin and development of blood vessels. Angiogenesis is the process through which new blood vessels are formed. Blood vessels (Fig. 1) are composed of two basic cell types: vascular endothelial cells and periendothelial cells (including…

A mineral with the chemical composition PbSO4. Anglesite occurs in white or gray, orthorhombic, tabular or prismatic crystals or compact masses (see illustration). It is a common secondary mineral, usually formed by the oxidation of galena. Fracture…

A mineral with the chemical composition CaSO4. Anhydrite occurs commonly in white and grayish granular masses, rarely in large, orthorhombic crystals (see illustration). Fracture is uneven and luster is pearly to vitreous. Hardness is 3–3.5 on Mohs…

The application of genetic principles for improving heredity for economically important traits in domestic animals. The science of animal breeding is based on fundamental genetic concepts. Through controlled mating and selection, which can involve…

A method used by animals to conceal themselves from an enemy by appearing to be a section of the natural background. Animal camouflage is a strategy that animal organisms use to avoid or deflect the attacks of predators by misleading the latter's…

The increase in size or weight of an animal. A number of processes are involved in growth. These include hyperplasia, or increase in cell number due to cell proliferation (cell division or mitosis) and recruitment from stem cell populations;…

The classification rank comprising the animals. Animals are taxonomically assigned to the classification kingdom known as Animalia. (The other kingdoms for classification of organisms include Plantae, Fungi, Protista, and Monera; in certain…

The formation of new individuals, which may occur by asexual or sexual means. All animals must reproduce asexually or sexually to create new individuals of their species. In asexual reproduction, a single parent can produce offspring that are…

The symmetrical disposition of organs and other constituent parts of the bodies of living animals with respect to imaginary axes. Animal symmetry relates the organization of parts in animal bodies to the geometrical design that each type suggests.…

The use of live nonhuman animals for scientific research and experimentation. Animal testing has been a cornerstone of biomedical research, having contributed greatly to the understanding of human biology, to the unraveling of the complex underlying…

An annual herb, Pimpinella anisum (family Apiaceae), whose fruit is used to provide food flavoring and to make a distilled oil. Anise is one of the earliest aromatics mentioned in literature. The anise plant (see illustration) is an annual herb that…

The carbonate mineral Ca(Fe,Mg)(CO3)2, also commonly containing some manganese. The mineral has hexagonal (rhombohedral) symmetry (see illustration) and has the cation-ordered structure of dolomite. The name is applied only to those species in which…

The calcium-rich plagioclase feldspar with composition Ab0An100 to Ab10An90 (Ab = NaAlSi3O8; An = CaAl2Si2O8), occurring in olivine-rich igneous rocks and rare volcanic ejecta (for example, at Mount Vesuvius, Italy, and Miyakejima, Japan). Hardness…

The name usually given to alkali feldspars which have a chemical composition ranging from Or40Ab60 to Or10Ab90± up to approximately 20 mole % An (Or, Ab, An = KAlSi3O8, NaAlSi3O8, CaAl2Si2O8) and which deviate in one way or another from monoclinic…

A rock composed of 90 vol % or more of plagioclase feldspar. Strictly, the rock is composed entirely of crystals discernible with the eye, but some finely crystalline examples from the Moon have been called anorthosite or anorthositic breccia. Two…

Oxygen-depleted regions in marine environments. The dynamic steady state between oxygen supply and consumption determines the oxygen concentration. In regions where the rate of consumption equals or exceeds the rate of supply, seawater becomes…

The Antarctic Circle is an imaginary line that delimits the northern boundary of Antarctica. It is a distinctive parallel of latitude at approximately 66°30′ south. Thus it is located about 4590 mi (7345 km) south of the Equator and about 1630 mi…

The Antarctic Ocean, sometimes called the Southern Ocean, is the watery belt surrounding Antarctica. It includes the great polar embayments of the Weddell Sea and Ross Sea, and the deep circumpolar belt of ocean between 50 and 60°S and the southern…

α Scorpii, a cool supergiant star of spectral type M1Ib, whose red color stands out in the midsummer sky. With an effective temperature of approximately 6000°F (3600 K), Antares resembles Betelgeuse, the brightest of the red supergiants, and would…

A magnesium-rich orthorhombic amphibole with perfect {210} cleavage and a color which varies from white to various shades of green and brown. It is a comparatively rare metamorphic mineral which occurs as slender prismatic needles, in fibrous…

An acute, infectious worldwide zoonotic disease (transmissible to humans) caused by the spore-forming bacterium Bacillus anthracis to which most animals, especially grazing herbivores, are susceptible. Anthrax is a serious, though rare, infectious…

An ongoing current event in which a large number of living species are threatened with extinction or are going extinct because of environmentally destructive human activities. The Earth is presently in the midst of a mass extinction event. In the…

The observation, measurement, and explanation of human variability in time and space. Anthropology is a science concerned with the interrelations of the biological, cultural, geographical, and historical aspects of humankind, including modern human…

The capability that a bacterium acquires to counteract an inhibitory chemical molecule or compound that was formerly effective in killing it or preventing its growth. Antibiotic resistance in bacteria (Fig. 1) is a hot topic in the media and in…

A fold in layered rocks in which the strata are inclined down and away from the axes. The simplest anticlines (see illustration) are symmetrical, but in more highly deformed regions they may be asymmetrical, overturned, or recumbent. Most anticlines…

Glycoprotein or protein molecules synthesized by polar and north temperate fishes to enable them to survive in freezing seawater. Similar antifreezes are found in some insects, but relatively little is known about their structure and…

A reaction that occurs when an antigen combines with a corresponding antibody to produce an immune complex. A substance that induces the immune system to form a corresponding antibody is called an immunogen. All immunogens are also antigens because…

A chemical substance used to destroy or prevent the growth of infectious microorganisms on or in a human or other animal body. An antiseptic is an antimicrobial compound or drug preparation that inhibits the spread and development of harmful…

An antibody that will combine with and generally neutralize a particular toxin. When the manifestations of a disease are caused primarily by a microbial toxin, the corresponding antitoxin, if available in time, may have a pronounced prophylactic or…

The main vessel of the systemic arterial circulation arising from the heart. The aorta (see illustration) is the main artery leaving the heart in all vertebrates and some invertebrates. In mammals, reptiles, and birds, which possess two heart…

One of the cellular aggregations traditionally known as chemoreceptors which are generally found adjacent to the aortic arch and pulmonary artery and which often are adjacent to the origin of the subclavian arteries (see illustration). The term is…

Apatite is a generic name used to describe calcium phosphates with the simplified formula Ca10(PO4)6X2 (X = F, Cl, OH). Three end members form a complete solid-solution series involving the halide and hydroxyl anions, and these are the species…

A fine-grained, sugary-textured rock, generally of granitic composition; also any body composed of such rock. GraniteThe light-colored rock consists chiefly of quartz, microcline, or orthoclase perthite and sodic plagioclase, with small amounts of…

A hydrous calcium potassium silicate containing fluorine. The composition is variable but approximates to KFCa4(Si2O5)4· 8H2O. It resembles the zeolites, with which it is sometimes classified, but differs from most zeolites in having no aluminum. It…

Programmed cell death triggered by extracellular signals or genetically determined events and carried out by physiological processes within the cell. Apoptosis is one of the major forms of cell death in multicellular organisms (necrosis is the other…