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The ongoing lower level of species extinction intensity occurring between episodes of mass extinctions. Extinction is a normal part of the history of life. Out of every 100 species that have ever lived on Earth, 99 are no longer in existence.…

Typically, an aquatic vertebrate with scales, gills, and paired fins. Fishes (Fig. 1) are found throughout the world's waterways, from saltwater oceans to freshwater streams and ponds. About 35,000 species of fishes have been described to date. Many…

A subdiscipline of paleobiology that investigates the processes of preservation and their influence on information in the fossil record. Taphonomy is the study of fossilization and considers the processes at work in the incorporation of living…

Most commonly, a generic name for all fossil resins, although it has been restricted by some to refer only to succinite, the mineralogical species of fossil resin that includes most of the Baltic Coast deposits. The time necessary to transform resin…

A genus of fossil birds in the extinct order Archaeopterygiformes, characterized by flight feathers like those of modern birds. Archaeopteryx is generally accepted as the oldest known fossil bird. It represents a unique snapshot of evolution; most…

A class in the phylum Mollusca that is exclusively marine and includes squids, octopuses, and cuttlefishes. The molluscan class Cephalopoda is one of the major taxonomic groupings of marine animals and contains both extant members—squids (Fig. 1),…

An order of the class Reptilia, including the turtles, terrapins, and tortoises; also termed Testudines. The order Chelonia (often referred to as Testudines) comprises one of the oldest groups of reptiles on Earth. Members of the Chelonia…

Any living or fossil member of the plant division Pinophyta, consisting chiefly of woody trees or shrubs that produce cones and evergreen needles. Conifers are predominantly cone-bearing, evergreen trees and shrubs belonging to the order Pinales, as…

Genealogists and anthropologists have long disputed the provenance of domestic dogs (Canis lupus familiaris). Although genetic analyses and fossil evidence indicate that the domestic dog descended from the gray wolf (Canis lupus), the exact location…

The death and disappearance of a species. The extinction of a species usually represents an end point in a long series of population extinctions. During the extinction process, unique evolutionary history is lost at every stage, but the death of the…

A record of earlier life buried in rock. Originally meaning any distinctive object that has been dug up (from Latin fodio, dig), the term “fossil” soon came to refer particularly to things resembling animals and plants. These objects were known…

The common name for a member of the agnathan fishes. The fish superclass Agnatha (subphylum Vertebrata, phylum Chordata) comprises the jawless vertebrates (agnathans). Members include cyclostomes (modern lampreys and hagfishes), various extinct…

A lineage of true elephants in the extinct genus Mammuthus that occupied Eurasia and North America in the late Cenozoic Era. Mammoths (genus Mammuthus) are extinct elephants assigned to the mammalian order Proboscidea (Fig. 1). They inhabited…

The widespread, catastrophic loss of many species in a relatively short period of geological time. Earth has supported life for approximately 3.5 billion years, and it is normal for species to become extinct and for new species to arise through…

A major phylum of the animal kingdom comprising a diverse group of unsegmented, soft-bodied invertebrates. The phylum Mollusca is one of the most significant groupings of invertebrate animals. Members of the Mollusca, termed mollusks or molluscs,…

A mammalian order containing the egg-laying platypus and echidnas. Members of the order Monotremata (monotremes) are egg-laying mammals belonging to the subclass Prototheria (in some alternative classifications, the monotremes have been assigned to…

A small extinct order of articulated brachiopods that ranges in age from Early to Middle Cambrian and includes the earliest known calcitic brachiopods. Classification The order Obolellida is included within the class Obolellata, subphylum…

An order of coleoid cephalopods, characterized by eight appendages (which encircle the mouth), a saclike body, and an internal shell that is much modified or reduced from that of its ancestors. Members of the order Octopoda (subclass Coleoidea,…

A morphologically distinct type of trace fossil left by boring organisms in hard biogenic substrates. Oichnus Bromley 1981 is a formal taxonomic name with the rank of ichnogenus (a trace fossil genus). Oichnus is a Latin-derived term that…

A suborder of trilobites (a class of extinct arthropods) and a diverse clade consisting of roughly 100 distinct species. The trilobites constituting the suborder Olenellina belong to the order Redlichiida in the class Trilobita. These extinct…

A class of the subphylum Asterozoa (phylum Echinodermata), comprising the brittle stars and basket stars, in which the arms are usually clearly demarcated from a central disk and perform whiplike locomotor movements, and the tube feet are…

A taxonomic fish group comprising the bony fishes. The Osteichthyes grouping (often considered a superclass) consists of the classes Actinopterygii (ray-finned fishes) and Sarcopterygii (lungfishes and lobefins, or coelacanths, but excluding…

Fossil lobe-finned (sarcopterygian) fishes of Devonian to Permian age (395–270 million years ago), the ancestral group for the land vertebrates, or “tetrapods.” Phylogeny The term “osteolepiforms” is applied to a range of fossil fishes, found…

An order of extinct jawless vertebrate fishes, also called Cephalaspida, known from the Middle Silurian to Upper Devonian of Europe, Asia, and North America. They were mostly small, about 2 in. (5 cm) to 2 ft (60 cm) in length. The head and part of…

A major taxon of the Crustacea containing small bivalved animals 0.004–1.4 in. (0.1–33 mm) long, with most between 0.04 and 0.08 in. (1 and 2 mm). Members of the Ostracoda inhabit aquatic environments in nearly all parts of the world.…

A popular name applied to several groups of extinct jawless vertebrates (fishes). Most ostracoderms were covered with an external skeleton or armor of bone, from which is derived their name, meaning “shell-skinned” (see illustration). They are known…

A peculiar arthropod, evidently related to the Pycnogonida and represented by a number of well-preserved fossils from the Devonian Hunsrück shales. Palaeoisopus was formerly considered by a number of paleontologists to have an anterior jointed…

A large, extinct assemblage of primitive ray-finned (actinopterygian) fishes known from the Silurian to Cretaceous, which were most numerous in Carboniferous and Permian times. They are known as fossils from all continents except Antarctica.…

A tiny fossil fish found only in rocks of Middle Devonian age within a small geographic region of Caithness, Scotland, and principally at one site, Achanarras Quarry, where it is surprisingly abundant. One of the most enigmatic of all fossil fishes,…

The study of chemical processes used by organisms that lived in the geological past. Most information on the nature of life in the geological past comes from the study of fossils; a record of biochemical processes that occurred can be found in the…

The study of pollen grains and spores, both extant and extinct, as well as other organic microfossils. Although the origin of the discipline dates back to the seventeenth century, when modern pollen was first examined microscopically, the term…

A fossil lobe-finned (sarcopterygian) fish from rock strata of the Baltic region dating to the Middle–Late Devonian Period (about 372–368 million years ago), which provides key evidence about the evolutionary transition from fish to land vertebrates…

An extinct order of relatively large placental mammals represented by Paleocene-Eocene fossils from western Europe, North America, and eastern Asia. Pantodonts were an early evolutionary experiment in large-bodied herbivory by primitive placental…

A medium-sized class of extinct blastozoan echinoderms containing about 17–19 genera, ranging from the late Early Ordovician to the Early Silurian throughout North America and in northern Europe. Their maximum standing diversity (13 genera) occurred…

An order of herbivorous, odd-toed, hoofed mammals, including the living horses, zebras, donkeys, rhinoceroses, tapirs, and their extinct relatives. Members of the order Perissodactyla, or perissodactyls, are odd-toed ungulates (mammals with hooves).…

A concentration of appreciable numbers of petrified tree trunks exposed at the earth surface, standing upright in the position of growth, lying prostrate, or both. Petrifaction is a form of mineral emplacement or embedding, by which the organic…

An order of mammals comprising the living pangolins and their poorly known fossil predecessors. All living members of the mammalian order Pholidota, that is, the pangolins (also termed scaly anteaters), belong to the family Manidae and are assigned…

An alleged fossil human based on craniodental fragments that were eventually discovered to constitute a skillful hoax. The Piltdown man hoax was the scientifically most successful fraud in the history of anthropology. Between 1908 and 1914, Charles…

One of the major divisions of the seed plants. The most familiar and common representatives of the division Pinophyta are the evergreen, cone-bearing trees of the order Pinales (within the class Pinopsida). Members are known colloquially as…

The single class of the division Pinophyta (Gymnospermae). The class Pinopsida contains both living and fossil taxa. The living representatives of the Pinopsida are woody plants; most are trees with a central axis and excurrent branches. Leaves are…

A class of fishes (Pisces) known from the Devonian Period, and a few that survived into the base of the Carboniferous. They were true fishes or gnathostomes, and can be distinguished from other fishes by the following characters: the gill chamber…

A small group of marine reptiles forming part of the Sauropterygia, known only from deposits of Triassic age in Europe, the Near East (Israel), North Africa (Tunisia), and China. The earliest occurrence of placodonts is in the uppermost Lower…

The worldwide array of plant life, including plants that have roots in the soil, plants that live on or within other plants and animals, plants that float on or swim in water, and plants that are carried in the air. Fungi used to be included in the…

A Paleozoic (Ordovician and Devonian) order of sea stars of the class Asteroidea; only two genera have been recognized. Platyasteridans have relatively elongate flat arms; broad, transversely aligned ventral arm plates; and paxilliform aboral plates…

An order of the Podocopa (Ostracoda) containing a single family, Cytherellidae, and two extinct superorders. Platycopids are small marine or brackish-water ostracods with an asymmetrical shell that is oblong, nearly rectangular, and laterally…

One of two Recent subclasses of Ostracoda. The two (Podocopa and Myodocopa) may share many characteristics but differ significantly in others. Shared characteristics include a carapace (shell) with a straight-to-concave ventral margin and somewhat…

The only order of the superorder Polymixiomorpha, comprising the beardfishes. The order Polymixiiformes (infraclass Teleostei, class Actinopterygii) consists of one family (Polymixiidae) with one genus (Polymixia) and ten extant species. Several…

A class within the phylum Mollusca whose members are popularly called chitons, coat-of-mail shells, or sea cradles. The class Polyplacophora (alternatively Loricata) lies within the phylum Mollusca and comprises the chitons. Members of classes…

A division of the plant kingdom, commonly called the ferns, which is widely distributed throughout the world but is most abundant and varied in moist, tropical regions. The Polypodiophyta are sometimes treated as a class Polypodiopsida of a broadly…

A distinctive and apparently ancient order of actinopterygian fishes comprising the subclass Cladistia, commonly known as bichirs and reedfishes. The following combination of characteristics distinguishes members of the order Polypteriformes from…

The fossil record of sharks (members of the subclass Elasmobranchii within the class Chondrichthyes, or cartilaginous fishes) dates back more than 420 million years. Over the course of this time, sharks have been among the top predators found in the…

An order of placental mammals containing the largest and most powerful living terrestrial mammals—the elephants. Proboscideans once had a wide distribution, reaching every continent except Australia and Antarctica. Although early forms lacked both…

One of the four subclasses of the class Mammalia. Prototheria contains a single order, the Monotremata. No ancestral genera of fossil monotremes are known, and the structure of the living monotremes is so specialized that the affinities of the…

A group long recognized as an order of fossil plants (subdivision Psilopsida) collected in rocks of Late Silurian and Devonian age. It has been subdivided, on the basis of many data, into three categories whose descriptions include the chief kinds…

An order of small, delicate insects, referred to as psocids, bark lice, or book lice. The order Psocoptera (an equivalent name is Corrodentia) belongs to a group (clade), the Acercaria, that also includes the Hemiptera (true bugs, aphids, and…

A subclass of the molluscan class Gastropoda, containing about 23,500 species of snails that are grouped into three orders (or superorders): Systellommatophora, Basommatophora, and Stylommatophora. Pulmonates include most of the common snails found…

An order of very specialized deep-bodied fishes near the holostean level of organization that are known only from the fossil record. They were a widespread group that first appeared in the Upper Triassic of Europe with the genus Eomesodon,…

A class of vertebrates composed of turtles, lizards, snakes, crocodilians, amphisbaenians, and tuataras, as well as a variety of extinct forms. The members of the vertebrate class Reptilia are tetrapods, meaning that they possess four limbs or are…

The most recent and successful group (infraclass) of actinopterygians (ray-finned fishes), first appearing in the Upper Triassic. The members of the Teleostei (teleosts) constitute one of the major fish groups belonging to the class Actinopterygii…

One of the largest recognized groups of early amphibians with about 180 described genera. The Temnospondyli first appear in the fossil record in the lower Carboniferous and reach substantial diversity in the later Carboniferous and Permian. The…

An investigation of 890-million-year-old limestone rocks excavated from the mountains of Canada's Northwest Territories has revealed possible evidence of the oldest animal life on Earth. Prior to this study, the oldest undisputed fossils of animal…

One of the four subclasses of the class Mammalia, including all living mammals except the monotremes. The Theria were by far the most successful of the several mammalian stocks that arose from the mammallike reptiles in the Triassic. The subclass is…