Nelsen, Olin E. Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania.
- Additional Reading
A tubuloalveolar or acinous skin gland, also known as the tear gland. The lacrimal glands develop from the skin epithelium which folds inward over the developing eye. Two types occur among the vertebrates, the lacrimal proper and the Harderian. These eye glands are first found in the amphibians, associated with the inside of the lower eyelid. In urodele amphibians the eye gland extends along the inner aspect of the lower eyelid. In Salamandra it becomes divisible into an anterior Harderian gland associated with lower eyelid structures and a posterior lacrimal gland below the upper eyelid. In frogs and toads only the Harderian gland is present and is associated with the nictitating membrane or third eyelid which develops in relation to the lower lid.
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