Van Fleet, Dick S. Department of Botany, University of Georgia, Athens, Georgia.
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The outermost cell layer of the cortex of plants. It forms a prominent layer immediately under the epidermis in many but not all plants (see illustration). In shoots, the hypodermis may be composed of parenchyma, collenchyma, or sclerenchyma and be from one to several cells thick. In roots, the hypodermis is often called the exodermis; it resembles the endodermis, and it develops Casparian strips, suberin deposits, and cellulose deposits impregnated with phenolic or quinoidal substances. Thus the root hypodermis is similar to the endodermis in cell wall anatomy and in its reaction to histochemical tests. The hypodermis is the mirror image of the endodermis in appearance. In the endodermis, wall deposits develop from the inner tangential wall outward; in the hypodermis, they develop from the outer tangential wall inward. See also: Cell walls (plant)
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